我国人工林林下灌木层植物多样性空间变异及影响要素

doi:10.12302/j.issn.1000-2006.202103014

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南京林业大学学报（自然科学版） ›› 2022, Vol. 46 ›› Issue (4): 144-152.doi: 10.12302/j.issn.1000-2006.202103014

我国人工林林下灌木层植物多样性空间变异及影响要素

郭弘婷¹(), 纪小芳², 汪成³, 邹汉鲁³, 王丽艳⁴, 姜姜¹^,^*()

1.南京林业大学,南方现代林业协同创新中心,江苏省水土保持与生态修复重点实验室,南京林业大学林学院,江苏南京 210037
2.中国农业科学院农业资源与农业区划研究所,北京 100081
3.永丰县官山林场,江西吉安 331500
4.江西省林业科学院,江西南昌 330032

收稿日期:2021-03-05 修回日期:2021-07-12 出版日期:2022-07-30 发布日期:2022-08-01
通讯作者: 姜姜
基金资助:
国家重点研发计划(2017YFC0505500);国家重点研发计划(2017YFC0505502);国家自然科学基金项目(32071612);国家自然科学基金项目(41701225)

Spatial variability and influencing factors of plant diversity in the shrub layers of artificial forests in China

GUO Hongting¹(), JI Xiaofang², WANG Cheng³, ZOU Hanlu³, WANG Liyan⁴, JIANG Jiang¹^,^*()

1. Co-Innovation Center for Sustainable Forestry in Southern China, Jiangsu Province Key Laboratory of Soil and Water Conservation and Ecological Restoration, College of Forestry, Nanjing Forestry University, Nanjing 210037, China
2. Institute of Agricultural Resource and Regional Planning, Chinese Academy of Agricultural Sciences, Beijing 100081, China
3. Yongfeng County Guanshan Forest Farm, Ji’an 331500, China
4. Jiangxi Academy of Forestry, Nanchang 330032, China

Received:2021-03-05 Revised:2021-07-12 Online:2022-07-30 Published:2022-08-01
Contact: JIANG Jiang

摘要/Abstract

摘要：

【目的】灌木层是森林生态系统的重要组成部分,建立我国人工林林下灌木层植物多样性数据库,为森林群落结构维持、生物多样性保护提供技术支撑。【方法】采用结构方程模型(SEM)揭示我国人工林林下灌木层植物多样性空间变异和影响要素。【结果】①林下灌木层植物多样性从南到北逐渐减小;②随着乔木胸径的增大,林下灌木层植物多样性逐渐增大,但分布均匀度有降低趋势;③随着人工林郁闭度的增加,植物多样性呈现先增大后减小趋势,郁闭度约为65.8%时Shannon-Wiener多样性指数达到峰值;④随着林分密度的增大,林下灌木层植物多样性逐渐减小;⑤温度、降水、郁闭度、林分胸径、树高和林分密度因子均对林下灌木层植物多样性具有直接或间接的作用,其中郁闭度对其影响存在阈值(65.8%),当郁闭度超过此阈值,温度和降水为主导因子,而郁闭度低于此阈值,温度的影响权重降低,林下光照的主导作用增强。【结论】在探究人工林林下灌木层植物多样性空间变异及影响要素的过程中,需要综合考虑各种影响因子,并借助多种多样性指数,综合评判林下植物多样性变化情况。本研究结果可为调控人工林林下灌木层植物多样性及林下植物恢复提供参考依据。

关键词: 人工林, 林下灌木层, 生物多样性, 结构方程模型(SEM)

Abstract:

【Objective】 The shrub layer is an important part of the forest ecosystem and plays a key role in maintaining the structure of the forest community and protecting biodiversity. The spatial variability and influencing factors of plant diversity in the undergrowth shrub layers of artificial forests in China were analyzed, providing a reference for regulation of plant diversity under plantations. 【Method】 In this paper, by establishing a database of plant diversity in undergrowth shrubs of artificial forests in China, structural equation modeling (SEM) was used to reveal the spatial variability and influencing factors of plant diversity in undergrowth shrubs in artificial forests. 【Result】 (1) Plant diversity in the forest shrub layer gradually decreased from south to north. (2) As the tree diameter at breast height increased, the plant diversity of the shrub layer gradually increased. However, the distribution uniformity tended to decrease. (3) As the canopy closure of the artificial forests increases, the plant diversity initially increased and then decreased. When the Shannon-Wiener diversity index reached its peak, the canopy closure was approximately 65.8%. (4) As the forest density increased, the plant diversity of the shrub layer gradually decreased. (5) Temperature, precipitation, canopy closure, diameter at breast height, tree height and stand density all had direct or indirect effects on the plant diversity of the understory shrub layer. The canopy closure had a threshold of 65.8%, and when it exceeded this threshold, temperature and precipitation were the dominant influencing factors. When the canopy closure was lower than this threshold, the weight of the influence of temperature decreased. The leading role of understory light was enhanced. 【Conclusion】 To explore the spatial variations and influencing factors of plant diversity in the understory shrub layer of artificial forests, it is necessary to consider various influencing factors, and evaluate the change in understory plant diversity with the help of the diversity index. The results of this study can provide a reference basis for regulating plant diversity in the understory shrub layer and the restoration of understory plants.

Key words: artificial forest, undergrowth shrub layer, biodiversity, structural equation model (SEM)

中图分类号:

S718.55+1.2

郭弘婷,纪小芳,汪成,等. 我国人工林林下灌木层植物多样性空间变异及影响要素[J]. 南京林业大学学报（自然科学版）, 2022, 46(4): 144-152.

GUO Hongting, JI Xiaofang, WANG Cheng, ZOU Hanlu, WANG Liyan, JIANG Jiang. Spatial variability and influencing factors of plant diversity in the shrub layers of artificial forests in China[J].Journal of Nanjing Forestry University (Natural Science Edition), 2022, 46(4): 144-152.DOI: 10.12302/j.issn.1000-2006.202103014.

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参考文献 35

[1]	张维伟, 薛文艳, 杨斌, 等. 桥山栎林群落结构特征与物种多样性相关关系分析[J]. 生态学报, 2019, 39(11): 3991-4001.
	ZHANG W W, XUE W Y, YANG B, et al. Differences in community characteristics,species diversity,and their coupling associations among three Quercus acutissima forests in Qiaoshan[J]. Acta Ecol Sin, 2019, 39(11):3991-4001.DOI:10.5846/stxb201804200901.
[2]	MO Q F, LIN Q, WANG W J, et al. Assessments of foliar functional traits of understory shrubs in two 13-year reforested plantations in subtropical China[J]. Glob Ecol Conserv, 2020, 23:e01202.DOI:10.1016/j.gecco.2020.e01202.
[3]	史作民, 刘世荣, 程瑞梅. 宝天曼地区栓皮栎林恢复过程中高等植物物种多样性变化[J]. 植物生态学报, 1998, 22(5): 3-5.
	SHI Z M, LIU S R, CHENG R M. Changes in plant species diversity in a restoration sequence of Quercus variabilis forest stands in Baotianman Mountain[J]. Chin Plant Ecol, 1998, 22(5):415-421.
[4]	杜忠, 蔡小虎, 包维楷, 等. 林下层植被对上层乔木的影响研究综述[J]. 应用生态学报, 2016, 27(3): 963-972.
	DU Z, CAI X H, BAO W K, et al. Understory effects on overstory trees: a review[J]. Chin J Appl Ecol, 2016, 27(3):963-972.DOI:10.13287/j.1001-9332.201603.033.
[5]	LI S F, HUANG X B, SHEN J Y, et al. Effects of plant diversity and soil properties on soil fungal community structure with secondary succession in the Pinus yunnanensis forest[J]. Geoderma, 2020, 379:114646.DOI:10.1016/j.geoderma.2020.114646.
[6]	YANG Y, ZHANG X Y, WANG H M, et al. How understory vegetation affects the catalytic properties of soil extracellular hydrolases in a Chinese fir (Cunninghamia lanceolata) forest[J]. Eur J Soil Biol, 2019, 90:15-21.DOI:10.1016/j.ejsobi.2018.11.004.
[7]	王云霖. 我国人工林发展研究[J]. 林业资源管理, 2019(1): 6-11.
	WANG Y L. Review on China’s plantation development since the reform and opening up[J]. For Resour Manag, 2019(1):6-11.DOI:10.13466/j.cnki.lyzygl.2019.01.002.
[8]	戴其林, 张超, 商克容, 等. 利用森林模拟优化模型(FSOS)分析森林经营单位合理年伐量[J]. 浙江农林大学学报, 2020, 37(5): 833-840.
	DAI Q L, ZHANG C, SHANG K R, et al. Rational annual cutting volume of forest management units based on FSOS[J]. J Zhejiang A&F Univ, 2020, 37(5):833-840.DOI:10.11833/j.issn.2095-0756.20190651.
[9]	夏丽丹, 于姣妲, 邓玲玲, 等. 杉木人工林地力衰退研究进展[J]. 世界林业研究, 2018, 31(2): 37-42.
	XIA L D, YU J D, DENG L L, et al. Researches on soil decline of Chinese fir plantation[J]. World For Res, 2018, 31(2):37-42.DOI:10.13348/j.cnki.sjlyyj.2017.0092.y.
[10]	王树梅, 庞元湘, 宋爱云, 等. 基于林龄的滨海盐碱地杨树刺槐混交土壤理化性质及草本植物多样性动态[J]. 生态学报, 2018, 38(18): 6539-6548.
	WANG S M, PANG Y X, SONG A Y, et al. Soil physiochemical properties and diversity of herbaceous plants dynamic on the different ages mixed forests of Populus × Euramercana ‘Neva’ and Robinia pseucdoacacia in coastal saline-alkali area[J]. Acta Ecol Sin, 2018, 38(18):6539-6548.DOI:10.5846/stxb201801220172.
[11]	李婷婷, 唐永彬, 周润惠, 等. 云顶山不同人工林林下植物多样性及其与土壤理化性质的关系[J]. 生态学报, 2020, 41(3): 1168-1177.
	LI T T, TANG Y B, ZHOU R H, et al. Understory plant diversity and its relationship with soil physicochemical properties in different plantations in Yunding Mountain[J]. Acta Ecol Sin, 2021, 41(3):1168-1177.DOI:10.5846/stxb202002270350.
[12]	黄超, 魏虹, 吴科君, 等. 马尾松林向香樟林改造林下植物功能多样性研究[J]. 生态学报, 2020, 40(13): 4573-4584.
	HUANG C, WEI H, WU K J, et al. The functional diversity of understory plants during the transformation from Pinus massoniana to Cinnamomum camphora forest[J]. Acta Ecol Sin, 2020, 40(13):4573-4584.DOI:10.5846/stxb201911152455.
[13]	王媚臻, 毕浩杰, 金锁, 等. 林分密度对云顶山柏木人工林林下物种多样性和土壤理化性质的影响[J]. 生态学报, 2019, 39(3): 981-988.
	WANG M Z, BI H J, JIN S, et al. Effects of stand density on understory species diversity and soil physicochemical properties of a Cupressus funebris plantation in Yunding Mountain[J]. Acta Ecol Sin, 2019, 39(3):981-988.DOI: 10.5846/stxb201803170528
[14]	闫东锋, 贺文, 杨喜田. 栓皮栎人工林灌木层植物多样性的空间分布及其与光环境的关系[J]. 应用生态学报, 2020, 31(11): 3605-3613.
	YAN D F, HE W, YANG X T. Spatial distribution of plant diversity in shrub layer of Quercus variabilis plantation and its relationship with light environment[J]. Chin J Appl Ecol, 2020, 31(11):3605-3613.DOI:10.13287/j.1001-9332.202011.002.
[15]	张柳桦, 齐锦秋, 李婷婷, 等. 林分密度对新津文峰山马尾松人工林林下物种多样性和生物量的影响[J]. 生态学报, 2019, 39(15): 5709-5717.
	ZHANG L H, QI J Q, LI T T, et al. Effects of stand density on understory plant diversity and biomass in a Pinus massoniana plantation in Wenfeng Mountain,Xinjin County[J]. Acta Ecol Sin, 2019, 39(15):5709-5717.DOI:10.5846/stxb201808151747.
[16]	李萌, 陈永康, 徐浩成, 等. 不同间伐强度对南亚热带杉木人工林林下植物功能群的影响[J]. 生态学报, 2020, 40(14): 4985-4993.
	LI M, CHEN Y K, XU H C, et al. Effects of different thinning intensities on undergrowth plant functional groups in subtropical Cunninghamia lanceolata plantations[J]. Acta Ecol Sin, 2020, 40(14):4985-4993.DOI:10.5846/stxb201905171015.
[17]	ZHANG C L, LI X W, CHEN Y Q, et al. Effects of Eucalyptus litter and roots on the establishment of native tree species in Eucalyptus plantations in South China[J]. For Ecol Manag, 2016, 375:76-83.DOI:10.1016/j.foreco.2016.05.013.
[18]	COOTE L, DIETZSCH A C, WILSON M W, et al. Testing indicators of biodiversity for plantation forests[J]. Ecol Indic, 2013, 32:107-115.DOI:10.1016/j.ecolind.2013.03.020.
[19]	陈露蔓, 吕倩, 刘思泽, 等. 柏木低效人工林开窗初期草本层植物多样性及生态位[J]. 应用与环境生物学报, 2021, 127(5):1178-1185.
	CHEN L M, LÜ Q, LIU S Z, et al. Diversity and niche characteristics of herbaceous layer plants in the early stage of forest gap reconstruction of inefficient Cupressus funebris plantations[J]. Chin J Appl Environ Biol, 2021, 27(5):1178-1185.DOI:10.19675/j.cnki.1006-687x.2020.05025.
[20]	曹小玉, 李际平, 赵文菲, 等. 基于结构方程模型分析林分空间结构对草本物种多样性的影响[J]. 生态学报, 2020, 40(24): 9164-9173.
	CAO X Y, LI J P, ZHAO W F, et al. Effects of stand spatial structure on herbaceous species diversity in forests based on structural equation modeling[J]. Acta Ecol Sin, 2020, 40(24):9164-9173.DOI:10.5846/stxb202001280179.
[21]	胡文浩, 张晓婧, 陈雅杰, 等. 坝上地区不同年代退耕还林生境的草本层植物多样性及影响因子[J]. 生态学报, 2021(3): 1-11.
	HU W H, ZHANG X J, CHEN Y J, et al. Herb diversity and its impact factors in different periods from 1970s to 2000s of the returning farmland to forest project in the Bashang area of north China[J]. Acta Ecol Sin, 2021, 41(3):1116-1126.DOI:10.5846/stxb201912042631.
[22]	方怡然, 李洁, 薛立. 不同林分改造模式对加勒比松林林下植物多样性和土壤物理性质的影响[J]. 中南林业科技大学学报, 2018, 38(6): 109-114.
	FANG Y R, LI J, XUE L. Impact of stand transformation on plant diversity and soil physical properties in Pinus caribaea stands[J]. J Central South Univ For Technol, 2018, 38(6):109-114.DOI:10.14067/j.cnki.1673-923x.2018.06.017.
[23]	周新. 基于结构方程模型的中国碳排放影响因素分析[D]. 北京: 北京交通大学, 2019.
	ZHOU X. Analysis of influence factors of China’s carbon emissions based on structural equation model[D]. Beijing: Beijing Jiaotong University, 2019.
[24]	王炎, 齐麟, 周莉, 等. 兴安落叶松(larix gmelinii)火后种子萌发影响因子研究[J]. 生态学报, 2021, 41(7): 2835-2844.
	WANG Y, QI L, ZHOU L, et al. A study on the influencing factors of Larix gmelinii post-fire seed germination[J]. Acta Ecol Sin, 2021, 41(7):2835-2844.DOI:10.5846/stxb201911252558.
[25]	张兴航, 张百平, 王晶, 等. 中国南北过渡带东段样带植被序列与气候分界问题[J]. 地理学报, 2021, 76(1): 30-43.
	ZHANG X H, ZHANG B P, WANG J, et al. North-south transect series in vegetation of Eastern China’s north-south transitional zone and their significance for determining climate dividing line[J]. Acta Geogr Sin, 2021, 76(1):30-43.DOI:10.11821/dlxb202101003.
[26]	孙启越, 张卫强, 赵连清, 等. 皆伐前后油松人工林下植物多样性变化及与土壤水分的关系[J]. 中南林业科技大学学报, 2020, 40(3): 119-129.
	SUN Q Y, ZHANG W Q, ZHAO L Q, et al. Variation of understory plant diversity and its relationship with soil moisture in Pinus tabulaeformis plantation before and after clearcutting[J]. J Central South Univ For Technol, 2020, 40(3):119-129.DOI:10.14067/j.cnki.1673-923x.2020.03.015.
[27]	吴世祥, 何聪, 杨丹, 等. 2009—2018年谢家店震后滑坡体植被恢复研究[J]. 水土保持研究, 2019, 26(6): 235-239,248.
	WU S X, HE C, YANG D, et al. Study of vegetation restoration after the earthquake on Xiejiadian landslide in the period 2009-2018[J]. Res Soil Water Conserv, 2019, 26(6):235-239,248.DOI:10.13869/j.cnki.rswc.20190831.001.
[28]	尤业明, 徐佳玉, 蔡道雄, 等. 广西凭祥不同年龄红椎林林下植物物种多样性及其环境解释[J]. 生态学报, 2016, 36(1): 164-172.
	YOU Y M, XU J Y, CAI D X, et al. Environmental factors affecting plant species diversity of understory plant communities in a Castanopsis hystrix plantation chronosequence in Pingxiang,Guangxi,China[J]. Acta Ecol Sin, 2016, 36(1):164-172.DOI:10.5846/stxb201411242332.
[29]	赵燕波, 张丹桔, 张健, 等. 不同郁闭度马尾松人工林林下植物多样性[J]. 应用与环境生物学报, 2016, 22(6): 1048-1054.
	ZHAO Y B, ZHANG D J, ZHANG J, et al. Understory vegetation diversity of Pinus massoniana plantations with various canopy density[J]. Chin J Appl Environ Biol, 2016, 22(6):1048-1054.DOI:10.3724/SP.J.1145.2016.04052.
[30]	贾亚运, 周丽丽, 吴鹏飞, 等. 不同发育阶段杉木人工林林下植被的多样性[J]. 森林与环境学报, 2016, 36(1): 36-41.
	JIA Y Y, ZHOU L L, WU P F, et al. Characterization of understory community of Cunninghamia lanceolata plantations at different developmental stages[J]. J For Environ, 2016, 36(1):36-41.DOI:10.13324/j.cnki.jfcf.2016.01.006.
[31]	王玲. 林分密度对油松人工林群落结构和植物多样性的影响[J]. 生态环境学报, 2020, 29(12): 2328-2336.
	WANG L. Effects of different stand densities on community structure and species diversity of Pinus tabulaeformis plantation[J]. Ecol Environ Sci, 2020, 29(12):2328-2336.DOI:10.16258/j.cnki.1674-5906.2020.12.003.
[32]	温晶, 张秋良, 李嘉悦, 等. 间伐强度对兴安落叶松林林下植被多样性及生物量的影响[J]. 中南林业科技大学学报, 2019, 39(5): 95-100, 118.
	WEN J, ZHANG Q L, LI J Y, et al. Effects of thinning intensity on diversity of undergrowth vegetation and biomass in Larix gmelini forest[J]. J Central South Univ For Technol, 2019, 39(5):95-100,118.DOI:10.14067/j.cnki.1673-923x.2019.05.014.
[33]	孙千惠, 吴霞, 王媚臻, 等. 林分密度对马尾松林林下物种多样性和土壤理化性质的影响[J]. 应用生态学报, 2018(3): 732-738.
	SUN Q H, WU X, WANG M Z, et al. Effects of stand density on understory species diversity and soil physicochemical properties of Pinus massoniana plantation[J]. Chin J Appl Ecol, 2018, 29(3):732-738.DOI:10.13287/j.1001-9332.201803.004.
[34]	丁继伟, 张芸香, 郭跃东, 等. 华北落叶松天然林密度对林下植被物种组成和多样性的影响研究[J]. 西北林学院学报, 2018, 33(4): 10-16.
	DING J W, ZHANG Y X, GUO Y D, et al. Influences of the density on understory species composition and diversity of Larix principis-rupprechtii natural forest[J]. J Northwest For Univ, 2018, 33(4):10-16.DOI:10.3969/j.issn.1001-7461.2018.04.02.
[35]	WULF M, NAAF T. Herb layer response to broadleaf tree species with different leaf litter[J]. Journal of Vegetation Science, 2009, 20(3): 517-526. DOI:10.1111/j.1654-1103.2009.05713.x.

我国人工林林下灌木层植物多样性空间变异及影响要素

Spatial variability and influencing factors of plant diversity in the shrub layers of artificial forests in China

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