38年生马尾松种源生长及材性联合分析

doi:10.12302/j.issn.1000-2006.202104044

国家林草科技领军期刊
中国精品科技期刊
中国高校百佳科技期刊
江苏省新闻出版政府奖期刊奖
RCCSE林学权威期刊（A+）
CSCD核心期刊
Scopus数据库收录期刊
中文核心期刊
SCD核心期刊

作者加群：102861116

微信公众号：南京林业大学学报

高级检索

南京林业大学学报（自然科学版） ›› 2022, Vol. 46 ›› Issue (3): 203-212.doi: 10.12302/j.issn.1000-2006.202104044

38年生马尾松种源生长及材性联合分析

胡兴峰¹(), 吴帆¹, 孙晓波¹, 陈厚平², 殷安政², 季孔庶¹^,^*()

1.南京林业大学林木遗传与生物技术省部共建教育部重点实验室,南方现代林业协同创新中心,江苏南京 210037
2.安徽省六安市裕安区国有林木良种场,安徽六安 237010

收稿日期:2021-04-26 接受日期:2021-11-08 出版日期:2022-05-30 发布日期:2022-06-10
通讯作者: 季孔庶
基金资助:
国家重点研发计划(2017YFD0600304);江苏高校优势学科建设工程资助项目(PAPD)

Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances

HU Xingfeng¹(), WU Fan¹, SUN Xiaobo¹, CHEN Houping², YIN Anzheng², JI Kongshu¹^,^*()

1. Key Laboratory of Forest Genetics & Biotechnology of Ministry of Education, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing 210037, China
2. State-owned Forest Tree Breeding Farm, Yu’an District, Lu’an City, Anhui Province, Lu’an 237010, China

Received:2021-04-26 Accepted:2021-11-08 Online:2022-05-30 Published:2022-06-10
Contact: JI Kongshu

摘要/Abstract

摘要：

【目的】通过分析已达成熟期的马尾松种源生长与材性变异,揭示不同性状在地理种源上的变异规律,并筛选适合在安徽省六安市及类似立地区域推广的速生且材质优良的纸浆材种源。【方法】以安徽省六安市裕安区国有林木良种场的38年生(成熟林)马尾松55个种源为试材,测定其胸径、树高、管胞长度及宽度、基本密度和微纤丝角,进行生长与材性联合分析。【结果】胸径、树高、材积、管胞长度、管胞宽度、微纤丝角和基本密度在种源水平上均差异极显著(P<0.01)。各性状表型变异系数、遗传变异系数分别为3.02%~58.61%、5.56%~28.16%;胸径、树高、材积、管胞长度、管胞宽度、基本密度和微纤丝角种源遗传力分别为0.665 7、0.558 9、0.616 9、0.996 3、0.965 6、0.739 3和0.823 3。相关性研究表明:生长性状与纬度呈显著负相关,与经度相关不显著;材性与经纬度均相关不显著,但表现出局部的变异规律;生长性状间相关性高于生长与材性间相关性;微纤丝角从髓心至树皮总体随着龄级的增大而减小,木材基本密度、管胞长和宽从髓心至树皮总体随着龄级的逐渐增大而增大,变化幅度随着龄级的增大而减小。通过综合指数选择出5个综合性状优良种源,其胸径、树高、材积、管胞长度与宽度、基本密度和微纤丝角平均遗传增益分别为3.69%、5.72%、15.12%、7.05%、-1.12%、0.17%和-6.46%。【结论】成熟林马尾松种源间差异显著,并且生长性状存在从南向北逐渐减小的变异规律,材性与经纬度不相关,对其进行纸浆材生长与材性联合选择具一定潜力,可为纸浆材马尾松良种选择提供可靠依据。

关键词: 马尾松, 种源, 基本密度, 管胞, 微纤丝角, 材性

Abstract:

【Objective】The growth provenance and wood property variation of mature masson pine (Pinus massoniana) are studied, revealing the variation rules for different traits at the level of geographical provenance, and screening suitable provenances for Lu’an City in Anhui Province and similar geographical areas.【Method】We selected the state-owned forest tree seed field in Yu’an District, Lu’an City, Anhui Province as the experimental site. 55 provenances were selected for 38-year-old (mature forest) P. massoniana as the test material. The tree height, diameter at breast height(DBH) volume, length and width of tracheid, basic density and microfibril angle are combined to analyze the growth and material properties.【Result】It was found that the diameter at breast height, tree height, volume, tracheid length, tracheid width, microfibril angle and basic density all showed highly significant differences at the provenance level (P < 0.01). The variation range of the phenotypic coefficients of variation and the genetic variation coefficients of each trait are 3.02%-58.61% and 5.56%-28.16%, respectively. The provenance heritability of the DBH, tree height, volume, tracheid length, tracheid width, basic density and microfibril angle are 0.665 7, 0.558 9, 0.616 9, 0.996 3, 0.965 6, 0.739 3 and 0.823 3, respectively. Growth traits are significantly negatively correlated with latitude, and are not significantly different from longitude. Wood properties at different latitude and longitude are both insignificantly different, but show local variation. In the correlation study, it was found that the correlation between growth traits is higher than that between growth and wood properties. The microfibril angle from the medulla core to the bark decreases with increase in age. The basic wood density, tracheid length and width from the pith to the bark increases with the gradual increase in age class, and the range of change decreases with the increase in age class. Through the comprehensive index, five provenances were selected with excellent comprehensive traits. The average genetic gains of DBH, tree height, volume, tracheid length and width, basic density and microfibril angle were 3.69%, 5.72%, 15.12%, 7.05%,-1.12%, 0.17% and -6.46%.【Conclusion】There is abundant genetic variation among the provenances of P. massoniana in mature forests, and growth characteristics show a variation law that gradually decreases from south to north, with wood properties not being related to latitude and longitude. The joint selection of pulp material growth and material properties provides a reliable basis for the selection of fine varieties of pulp material in P. massoniana.

Key words: masson pine (Pinus massoniana), provenance, basic density, tracheid, microfibril angle, wood property

中图分类号:

S722

胡兴峰,吴帆,孙晓波,等. 38年生马尾松种源生长及材性联合分析[J]. 南京林业大学学报（自然科学版）, 2022, 46(3): 203-212.

HU Xingfeng, WU Fan, SUN Xiaobo, CHEN Houping, YIN Anzheng, JI Kongshu. Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances[J].Journal of Nanjing Forestry University (Natural Science Edition), 2022, 46(3): 203-212.DOI: 10.12302/j.issn.1000-2006.202104044.

图/表 11

表1

供试马尾松种源及编号"

种源编号 provenance No.	种源 provenance	东经/(°) longitude (E)	北纬(°) latitude(N)	种源编号 provenance No.	种源 provenance	东经/(°) longitude (E)	北纬(°) latitude(N)
1	陕西城固	107.353 3	33.157 1	33	贵州黎平	109.140 1	26.237 4
2	河南新县	114.891 8	31.650 2	34	四川蒲江	103.509 1	30.203 0
3	安徽霍山	116.337 2	31.399 6	35	浙江鄞县	121.553 1	29.823 7
4	浙江遂昌	119.278 4	28.600 0	36	江西崇义	114.313 4	25.686 6
5	江西安远	115.394 2	25.142 7	39	江西崇仁	116.081 8	27.757 5
6	湖南慈利	111.148 9	29.434 5	40	浙江三门	121.401 4	29.112 9
7	湖北通山	114.487 7	29.613 6	41	安徽太平	118.143 3	30.299 6
8	福建漳平	117.427 1	25.296 3	42	浙江开化	118.420 9	29.143 2
9	广东高川	110.853 7	21.923 0	43	江西石城	116.346 3	26.320 0
11	贵州黄平	107.919 5	26.911 3	44	福建古田	118.746 8	26.581 7
12	四川南江	106.831 0	32.353 0	45	广东信宜	110.952 4	22.360 6
13	湖南安化	111.217 8	28.380 6	46	广西忻城	108.671 4	24.071 4
14	浙江庆远	119.066 0	27.626 0	47	浙江嵊县	120.837 4	29.567 3
15	江西安福	114.622 5	27.399 6	48	江西万载	114.453 7	28.114 7
17	湖南临湘	113.458 0	29.482 8	49	陕西南郑	106.940 1	33.006 1
18	福建德化	118.240 6	25.498 9	50	河南鹤峰	114.303 2	35.754 0
19	广东广宁	112.445 1	23.642 6	51	安徽屯溪	118.320 3	29.703 1
20	贵州都匀	107.531 1	26.267 7	52	浙江民生	120.752 9	30.104 0
21	浙江缙云	120.091 7	28.665 7	53	江西清江	114.798 0	29.193 5
22	江西吉安	114.997 6	27.120 2	54	湖南资兴	113.239 1	25.982 4
25	贵州凯里	107.986 3	26.574 1	55	福建邵武	117.501 1	27.348 5
26	河南固始	115.658 8	32.174 7	58	广西茶城	110.251 2	24.490 7
27	安徽马头	118.509 3	30.788 3	59	贵州德江	108.124 1	28.272 5
28	浙江永康	120.052 8	28.894 2	60	四川涪陵	107.394 7	29.707 8
29	江西余江	116.860 2	28.206 2	61	湖南永顺	109.862 4	28.987 8
30	湖南江永	111.345 8	25.282 3	63	江西资溪	109.862 4	28.987 8
31	福建三明	117.648 9	26.270 3	64	浙江西溪	120.080 0	30.271 9
32	广东罗定	111.573 0	22.776 4

表1

图1

表2

表3

表4

表5

图2

表6

图3

表7

表8

参考文献 34

[1]	秦国峰, 周志春. 中国马尾松优良种质资源[M]. 北京: 中国林业出版社, 2012.
	QIN G F, ZHOU Z C. Germplasm resources of Chinese masson pine[M]. Beijing: China Forestry Publishing House, 2012.
[2]	谭健晖, 冯源恒, 贾婕, 等. 马尾松半同胞家系纤维形态遗传变异及纸浆材优良家系选择[J]. 南京林业大学学报(自然科学版), 2012, 36(6): 8-12.
	TAN J H, FENG Y H, JIA J, et al. Study on the genetic variation of wood fiber morphology in a half-sib mason pine family and the selection of fine paper pulp making family[J]. J Nanjing For Univ (Nat Sci Ed), 2012, 36(6): 8-12. DOI: 10.3969/j.issn.2095-1914.2012.03.017. doi: 10.3969/j.issn.2095-1914.2012.03.017
[3]	卢翠香, 项东云, 陈健波, 等. 人工林马尾松管胞形态特征及其变异[J]. 浙江农林大学学报, 2012, 29(4): 588-594.
	LU C X, XIANG D Y, CHEN J B, et al. Fiber characteristics and variation patterns of Pinus massoniana[J]. J Zhejiang A & F U, 2012, 29(4): 588-594. DOI: cnki:sun:zjlx.0.2012-04-018. doi: cnki:sun:zjlx.0
[4]	张波. 马尾松木材管胞形态及微力学性能研究[D]. 北京: 中国林业科学研究院, 2007.
	ZHANG B. Morphology and micromechanics of masson pine tracheids[D]. Beijing: Chinese Academy of Forestry, 2007.
[5]	黄振英. 马尾松正常木与应压木生长应力及材性的比较研究[D]. 合肥: 安徽农业大学, 2004.
	HUANG Z Y. Comparative study on the growth stresses and the properties of normal wood and compression wood in masson pine[D]. Hefei: Anhui Agricultural University, 2004.
[6]	CHEN L, JIANG Z H, ZHAO R J, et al. Study situation of properties and modification of masson pine in China[J]. Chinese Forestry Science and Technology, 2009, 8(1): 29-35.
[7]	庄尔奇, 刘强, 高方彬, 等. 湖北省马尾松产区区划研究[J]. 华中农业大学学报, 1997, 16(1): 81-87.
	ZHUANG E Q, LIU Q, GAO F B, et al. Studies on productive region division of Masson pine(Pinus massoniana) in Hubei Province[J]. J Huazhong Agri Univ, 1997, 16(1): 81-87.
[8]	江西省马尾松产区区划、立地分类与评价及经营数表编制协作组. 江西省马尾松产区区划立地分类与评价及经营数表编制(总报告)[J]. 江西农业大学学报, 1987, 9(3): 1-8.
	Division of Pinus massoniana Production Area, Site Classification and Evaluation, and Establishment of Management Data Table in Jiangxi Province. Masson pine of the division site classification and evaluation of the number of operators and Jiangxi prepared (general report)[J]. Acta Agriculture Universitatis Jiangxiensis, 1987, 9(3): 1-8.
[9]	杨宗武, 郑仁华, 傅忠华, 等. 马尾松工业用材优良家系选择的研究[J]. 林业科学, 2003, 39(S1): 74-80.
	YANG Z W, ZHENG R H, FU Z H, et al. Study on selection of excellent families for industrial wood of Pinus massoniana[J]. Sci Silvae Sin, 2003, 39(S1): 74-80.
[10]	徐慧兰, 谭健晖, 范歌, 等. 3个马尾松种源材性差异及相关性[J]. 广西林业科学, 2017, 46(1):17-20.
	XU H L, TAN J H, FAN G, et al. The differences and correlation in wood properties of three provenances of Pinus massoniana[J]. Guangxi For Sci, 2017, 46(1):17-20.DOI: 10.19692/j.cnki.gfs.2017.01.004. doi: 10.19692/j.cnki.gfs.2017.01.004
[11]	冯源恒, 杨章旗, 李火根, 等. 不同时期广西马尾松优良种源的遗传多样性变化趋势[J]. 南京林业大学学报(自然科学版), 2016, 40(5):41-46.
	FENG Y H, YANG Z Q, LI H G, et al. A study on changes of genetic diversity for nearly 50 years in superior provenances of Pinus massoniana in Guangxi[J]. J Nanjing For Univ (Nat Sci Ed), 2016, 40(5):41-46.DOI: 10.3969/j.issn.1000-2006.2016.05.007. doi: 10.3969/j.issn.1000-2006.2016.05.007
[12]	张明慧, 陈虎, 何长虹, 等. 马尾松改良代种子园不同家系苗期生长差异分析[J]. 广西林业科学, 2017, 46(1):21-26.
	ZHANG M H, CHEN H, HE C H, et al. Growth variance analysis of different families seedling in advanced seed orchard of Pinus massoniana[J]. Guangxi For Sci, 2017, 46(1):21-26.DOI: 10.19692/j.cnki.gfs.2017.01.005. doi: 10.19692/j.cnki.gfs.2017.01.005
[13]	陈虎, 张明慧, 卢开成, 等. 修剪强度对马尾松成年种子园母树的影响[J]. 福建林业科技, 2017, 44(1):38-42.
	CHEN H, ZHANG M H, LU K C, et al. Effect of different pruning intensity in the seed orchard of Pinus massoniana L.trees[J]. J Fujian For Sci Technol, 2017, 44(1):38-42.DOI: 10.13428/j.cnki.fjlk.2017.01.009. doi: 10.13428/j.cnki.fjlk.2017.01.009
[14]	吴晓刚, 陈佩珍, 韦蔷, 等. 马尾松PmCOMT基因的克隆及实时定量表达分析[J]. 分子植物育种, 2019, 17(3):789-795.
	WU X G, CHEN P Z, WEI Q, et al. Cloning and real-time quantitative expression analysis on PmCOMT gene from Pinus massoniana[J]. Mol Plant Breed, 2019, 17(3):789-795.DOI: 10.13271/j.mpb.017.000789. doi: 10.13271/j.mpb.017.000789
[15]	周紫晶, 范付华, 尚先文, 等. 马尾松大小径材miRNA的差异表达分析[J]. 植物生理学报, 2020, 56(10):2111-2120.
	ZHOU Z J, FAN F H, SHANG X W, et al. Differential expression analysis of miRNA between large diameter wood and small diameter wood of Pinus massoniana[J]. Plant Physiol J, 2020, 56(10):2111-2120.DOI: 10.13592/j.cnki.ppj.2020.0247. doi: 10.13592/j.cnki.ppj.2020.0247
[16]	毛桃. 马尾松优树子代测定林生长和材质的遗传分析及联合选择[D]. 南京: 南京林业大学, 2007.
	MAO T. Genetic analysis and combined selection for growth and wood quality of half-sib family of masson pine[D]. Nanjing: Nanjing Forestry University, 2007.
[17]	林景泉. 18年生马尾松种子园自由授粉子代速生优质家系选择[J]. 三明学院学报, 2015, 32(2):7-11.
	LIN J Q. Family selection of fast growth and high quality in the 18-year-old open pollination progenies of Pinus massoniana seed orcard[J]. J Sanming Univ, 2015, 32(2):7-11.DOI: 10.14098/j.cn35-1288/z.2015.02.002. doi: 10.14098/j.cn35-1288/z.2015.02.002
[18]	谭健晖, 黄永利, 冯源恒, 等. 马尾松家系木材管胞形态特征的年龄变异[J]. 西南林业大学学报, 2012, 32(3):80-83.
	TAN J H, HUANG Y L, FENG Y H, et al. Age variation of wood tracheid morphology of Pinus massoniana families[J]. J Southwest For Coll, 2012, 32(3):80-83.DOI: 10.3969/j.issn.2095-1914.2012.03.017. doi: 10.3969/j.issn.2095-1914.2012.03.017
[19]	张水生. 马尾松种源地理变异规律的初步研究[J]. 江西农业大学学报, 1999, 21(4):597-601.
	ZHANG S S. A preliminary study on the law of geographic variation on the Pinus massoniana[J]. Acta Agric Univ Jiangxiensis, 1999, 21(4):597-601.
[20]	林思京. 25年生马尾松生长和木材基本密度家系变异与选择[J]. 林业科学研究, 2010, 23(6):804-808.
	LIN S J. Growth and wood density of 25-year-old masson’s pine:inter-family variation and selection[J]. For Res, 2010, 23(6):804-808.DOI: 10.13275/j.cnki.lykxyj.2010.06.005. doi: 10.13275/j.cnki.lykxyj.2010.06.005
[21]	金国庆, 秦国峰, 刘伟宏, 等. 不同林龄马尾松的种源选择效果[J]. 林业科学, 2011, 47(2):39-45.
	JIN G Q, QIN G F, LIU W H, et al. Provenance selection effect at different stand age of Pinus massoniana[J]. Sci Silvae Sin, 2011, 47(2):39-45.
[22]	武紫娟. 马尾松半同胞子代测定林变异分析及优良家系选择[D]. 雅安: 四川农业大学, 2016.
	WU Z J. Variation analysis and superior families selection on half-sib progeny tested forests of Pinus massoniana[D]. Yaan: Sichuan Agricultural University, 2016.
[23]	林能庆. 闽西马尾松优树子代测定及优良单株选择[J]. 南京林业大学学报(自然科学版), 2013, 37(5):31-34.
	LIN N Q. Plus tree selection and progenies test of Pinus massoniana in western Fujian Province[J]. J Nanjing For Univ (Nat Sci Ed), 2013, 37(5):31-34.
[24]	季孔庶, 毛桃, 康建诚, 等. 马尾松材性遗传改良研究进展[J]. 林业科技开发, 2011, 25(5):1-6.
	JI K S, MAO T, KANG J C, et al. Research progress on genetic improvement of masson pine wood properties[J]. Journal of Forestry Engineering, 2011, 25(5): 1-6.DOI: 10.3969/j.issn.1000-8101.2011.05.001. doi: 10.3969/j.issn.1000-8101.2011.05.001
[25]	SMITH D M. Maximum moisture content method for determining specific gravity of small wood samples:rept. No. 2014[J]. Wisconsin:Forest Products Laboratory, Forest Service, US Department of Agriculture, 1954.
[26]	李光友, 徐建民, 陆钊华, 等. 尾叶桉二代测定林家系的综合评选[J]. 林业科学研究, 2005, 18(1):57-61.
	LI G Y, XU J M, LU Z H, et al. Studies on index selections of Eucalyptus urophylla families[J]. For Res, 2005, 18(1):57-61.DOI: 10.3321/j.issn:1001-1498.2005.01.011. doi: 10.3321/j.issn:1001-1498.2005.01.011
[27]	孙戴妍, 尚旭岚, 洑香香, 等. 青钱柳胸径生长和木材密度的地理变异规律[J]. 南京林业大学学报(自然科学版), 2017, 41(4):1-5.
	SUN D Y, SHANG X L, FU X X, et al. Regularity on geographic variation in DBH growth and wood density of Cyclocarya paliurus[J]. J Nanjing For Univ (Nat Sci Ed), 2017, 41(4):1-5.DOI: 10.3969/j.issn.1000-2006.201610005. doi: 10.3969/j.issn.1000-2006.201610005
[28]	刘青华. 马尾松生长与材性的遗传变异、基因作用方式及环境影响[D]. 北京: 中国林业科学研究院, 2010.
	LIU Q H. Genetic variation,gene control pattern and environment effect of growth and wood property in Pinus massoniana[D]. Beijing: Chinese Academy of Forestry, 2010.
[29]	骆秀琴, 姜笑梅, 殷亚方, 等. 人工林马尾松木材性质的变异[J]. 林业科学研究, 2002, 15(1):28-33.
	LUO X Q, JIANG X M, YIN Y F, et al. Variations in wood properties of masson pine (Pinus massoniana L.) plantation[J]. For Res, 2002, 15(1):28-33.DOI: 10.3321/j.issn:1001-1498.2002.01.005. doi: 10.3321/j.issn:1001-1498.2002.01.005
[30]	周亮, 刘盛全, 田根林, 等. 欧美杨107杨木材解剖特性的径向变异[J]. 东北林业大学学报, 2010, 38(10):1-4.
	ZHOU L, LIU S Q, TIAN G L, et al. Radial variation of anatomical properties of poplar clone 107(Populus × euramericana cv.‘Neva’)[J]. J Northeast For Univ, 2010, 38(10):1-4.DOI: 10.13759/j.cnki.dlxb.2010.10.023. doi: 10.13759/j.cnki.dlxb.2010.10.023
[31]	郑仁华, 陈国金, 俞白楠, 等. 马尾松家系木材基本密度遗传变异的研究[J]. 西北林学院学报, 2001, 16(4):6-9.
	ZHENG R H, CHEN G J, YU B N, et al. On genetic variations of wood basic density in masson pine family[J]. J Northwest For Univ, 2001, 16(4):6-9.DOI: 10.3969/j.issn.1001-7461.2001.04.002. doi: 10.3969/j.issn.1001-7461.2001.04.002
[32]	徐立安, 陈天华, 王章荣, 等. 马尾松种源子代材性变异与制浆造纸材优良种源选择[J]. 南京林业大学学报, 1997, 21(2):1-6.
	XU L, CHEN T H, WANG Z R, et al. Variation of provenance progenies in wood property and provenance selection for pulpwood of masson pine[J]. J Nanjing For Univ, 1997, 21(2):1-6.
[33]	范林元, 赖焕林, 季孔庶, 等. 马尾松实生种子园建园家系遗传值估算与优良家系评选[J]. 东北林业大学学报, 2004, 32(4):3-5,11.
	FAN L Y, LAI H L, JI K S, et al. Breeding value estimation and family selection for seedling seed orchard of masson’s pine[J]. J Northeast For Univ, 2004, 32(4):3-5,11.DOI: 10.3969/j.issn.1000-5382.2004.04.002. doi: 10.3969/j.issn.1000-5382.2004.04.002
[34]	王明庥. 林木育种学概论[M]. 北京: 中国林业出版社, 1989.

性状 trait	均值 mean	标准差 SD	表型变异系数/% phenotypic coefficient of variation
DBH/cm	19.40	5.305 8	27.49
H/m	16.470 1	2.380 0	14.54
V/m³	0.251 1	0.144 0	58.61
TL/μm	3285.60	99.773 4	3.02
TW/μm	43.87	2.073 7	4.95
BD/(g·cm^-3)	0.568 9	0.033 9	5.93
MFA/(°)	29.23	1.654 7	5.70

性状 trait	变异来源 source of variation	自由度 df	均方 MS	F	P	显著性 signi- ficance
DBH	区组效应	5	418.930 6	30.366 4	1.82×10^-24	**
	种源效应	54	41.264 3	2.991 1	2.64×10⁹	**
H	区组效应	5	48.503 1	13.739 2	5.85×10^-12	**
	种源效应	54	8.004 2	2.267 3	9.92×10^-6	**
V	区组效应	5	0.243 5	19.966 7	6.18×10^-17	**
	种源效应	54	0.031 8	2.610 4	2.12×10^-7	**
TL	区组效应	5	48 078.271 4	24.986 3	1.10×10	**
	种源效应	54	525 894.681 6	273.307 2	9.81×10^-207	**
TW	区组效应	5	1.474 1	1.118 9	3.51×10^-1
	种源效应	54	38.298 2	29.070 7	3.60×10^-85	**
BD	区组效应	5	0.002 6	1.949 1	5.14×10^-2
	种源效应	54	0.005 2	3.836 4	2.64×10^-15	**
MFA	区组效应	5	18.105 9	5.407 1	5.78×10³	**
	种源效应	54	18.952 9	5.660 0	1.13×10^-14	**

性状 trait	方差variance						变异系数/% CV			种源遗传力 provenance heritability
性状 trait	表型 phenotypic	环境 environmental		遗传 genetic		表型 phenotypic		遗传 genetic		种源遗传力 provenance heritability
DBH	28.151 5		9.7090		1.381 0		27.49		7.13	0.665 7
H	5.664 4		0.8680		6.282 0		14.54		12.92	0.558 9
V	0.020 7		0.005 0		0.005 0		58.61		28.16	0.616 9
TL	9 954.731 3		874.155 0		87 648.621 0		3.02		9.01	0.996 3
TW	4.725 0		0.025 0		6.392 0		4.95		5.76	0.965 6
BD	0.001 1		0.000 048		0.001 0		5.93		5.56	0.739 3
MFA	2.738 0		0.329 0		6.336 0		5.70		8.60	0.823 3

项目item	H	DBH	V	TL	TW	BD	MFA
经度longitude(E)	-0.014(0.920)	-0.070(0.612)	-0.036(0.794)	-0.006(0.967)	-0.040(0.770)	0.094(0.494)	-0.072(0.599)
纬度latitude(N)	-0.427^**(0.001)	-0.561^**(<0.01)	-0.539^**(<0.01)	-0.071(0.606)	-0.140(0.308)	0.208(0.128)	0.011(0.939)

性状 trait	自由度 df	均方 MS	F	P	显著性 sig
TL	7	4 322.129 0	313.344 0	1.08×10^-274	***
TW	7	3.746 0	825.129 0	P<0.01	***
BD	7	1 012 684 377.190 0	6 467.518 0	P<0.01	***
MFA	7	16 358.522 0	391.205 0	P<0.01	***

38年生马尾松种源生长及材性联合分析

Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances

RichHTML

PDF

可视化

摘要/Abstract

引用本文

使用本文

图/表 11

参考文献 34

相关文章 15

编辑推荐

Metrics

本文评价

作者

读者

审稿

性状trait	DBH	H	V	TL	TW	BD	MFA
DBH	—	0.635^**	0.963^**	-0.016	0.018	-0.231^**	0.010
		(<0.01)	(<0.01)	(0.624)	(0.575)	(<0.01)	(0.899)
H	0.632^**	—	0.719^**	-0.008	0.023	-0.055	0.024
	(<0.01)		(<0.01)	(0.794)	(0.477)	(0.225)	(0.758)
V	0.735^**	0.967^**	—	0	0.025	-0.197^**	-0.008
	(<0.01)	(<0.01)		(0.994)	(0.438)	(<0.01)	(0.915)
TL	0.047	0.091	0.124	—	0.375^**	0.054	-0.098
	(0.733)	(0.507)	(0.369)		(<0.01)	(0.229)	(0.211)
TW	0.203	0.147	0.204	0.524^**	—	-0.029	0.027
	(0.137)	(0.284)	(0.135)	(<0.01)		(0.520)	(0.734)
BD	-0.417^**	-0.581^**	-0.516^**	0.120	-0.087	—	0.036
	(0.002)	(<0.01)	(<0.01)	(0.385)	(0.526)		(0.645)
MFA	-0.333^*	-0.121	-0.155	-0.098	-0.028	0.033	—
	(0.013)	(0.377)	(0.259)	(0.478)	(0.839)	(0.811)

种源号 provenance No.	H/m	DBH/cm	V/m³	TL/μm	TW/μm	BD/(g·cm^-3)	MFA/(°)	I
5	18.93	22.01	0.355 1	3 644.5	46.9	0.601 6	24.8	9.342
8	17.28	22.11	0.344 2	3 399.7	42.9	0.591 6	29.4	8.042
13	17.96	19.93	0.286 9	3 470.8	42.3	0.558 4	24.8	8.584
40	16.18	18.78	0.233 4	3 403.9	41.8	0.569 8	24.7	8.057
58	16.55	24.10	0.343 6	3 672.5	42.9	0.529 6	31.1	7.753
总体均值 population mean	16.47	19.40	0.251 1	3 285.6	43.9	0.568 9	29.3	—
入选群体均值 selected group mean	17.38	21.39	0.312 7	3 518.3	43.4	0.570 2	27.0	—
遗传增益/% genetic gain	3.69	5.72	15.12	7.05	-1.12	0.17	-6.46	—

[1]	张瑞, 周正虎, 王传宽, 金鹰. 东北温带森林不同材性树种木质部解剖和水力性状[J]. 南京林业大学学报（自然科学版）, 2024, 48(3): 229-236.
[2]	武燕, 黄青, 刘讯, 郑睿, 岑佳宝, 丁波, 张运林, 符裕红. 西南喀斯特地区马尾松人工林林龄对土壤理化性质的影响[J]. 南京林业大学学报（自然科学版）, 2024, 48(3): 99-107.
[3]	范明阳, 胡萌, 杨园, 方炎明. 中国东部地区马尾松与黄山松群落分类及群落结构和物种多样性特征[J]. 南京林业大学学报（自然科学版）, 2024, 48(1): 47-58.
[4]	王章荣, 季孔庶, 徐立安, 邹秉章, 林能庆, 林景泉. 马尾松实生种子园营建技术、现实增益及多世代低成本经营新模式探讨[J]. 南京林业大学学报（自然科学版）, 2023, 47(6): 9-16.
[5]	王宇, 易艳灵, 刘海, 文晓晨, 李天一, 尹海锋, 李贤伟, 范川. 两种采伐方式对马尾松人工林林分空间结构的影响[J]. 南京林业大学学报（自然科学版）, 2023, 47(5): 138-146.
[6]	盖军鹏, 陈东升, 贾炜玮, 王政. 基于种源和气候效应的日本落叶松树高生长模型研究[J]. 南京林业大学学报（自然科学版）, 2023, 47(4): 51-60.
[7]	孙荣喜, 潘昕昊, 仲小茹, 李桂盛. 不同种源米槠种子形态特征与营养成分变异分析[J]. 南京林业大学学报（自然科学版）, 2023, 47(2): 27-34.
[8]	竹磊, 徐军亮, 章异平, 罗鹏飞, 师志强, 候佳玉, 翟乐鑫. 河南洛阳马尾松树干液流昼夜变化特征及其影响因子分析[J]. 南京林业大学学报（自然科学版）, 2023, 47(1): 92-100.
[9]	孙薇, 王斌, 楚秀丽, 王秀花, 张东北, 吴小林, 周志春. 马尾松容器苗生长和养分性状对磷添加和接种菌根菌的响应及关联[J]. 南京林业大学学报（自然科学版）, 2023, 47(1): 226-233.
[10]	季孔庶, 徐立安, 王登宝, 倪州献, 王章荣. 中国马尾松遗传改良研究历程与成就[J]. 南京林业大学学报（自然科学版）, 2022, 46(6): 10-22.
[11]	王立超, 苏胜荣, 陈凤毛, 董晓燕, 田成连, 王洋. 黄山马尾松林天牛及携带线虫种类初步调查[J]. 南京林业大学学报（自然科学版）, 2022, 46(4): 29-35.
[12]	吴佳雯, 尹艳楠, 谈家金, 郝德君. 蜡样芽孢杆菌NJSZ-13菌株诱导马尾松抗松材线虫病研究[J]. 南京林业大学学报（自然科学版）, 2022, 46(4): 53-58.
[13]	吴帆, 朱沛煌, 季孔庶. 马尾松分布格局对未来气候变化的响应[J]. 南京林业大学学报（自然科学版）, 2022, 46(2): 196-204.
[14]	高景斌, 徐六一, 叶建仁. 马尾松松材线虫病抗性无性系的筛选和遗传多样性分析[J]. 南京林业大学学报（自然科学版）, 2021, 45(5): 109-118.
[15]	尹艳楠, 谈家金, 李梦伟, 许嘉麟, 郝德君. 蜡样芽孢杆菌NJSZ-13菌株防治松材线虫病研究[J]. 南京林业大学学报（自然科学版）, 2021, 45(3): 152-158.