南京林业大学学报(自然科学版) ›› 2024, Vol. 48 ›› Issue (1): 47-58.doi: 10.12302/j.issn.1000-2006.202210029
所属专题: 森林生态系统生物多样性研究专题
• 专题报道Ⅱ:森林生态系统生物多样性研究专题(执行主编 薛建辉 方炎明) • 上一篇 下一篇
收稿日期:
2022-10-23
修回日期:
2023-01-25
出版日期:
2024-01-30
发布日期:
2024-01-24
通讯作者:
方炎明
基金资助:
FAN Mingyang(), HU Meng, YNAG Yuan, FANG Yanming()
Received:
2022-10-23
Revised:
2023-01-25
Online:
2024-01-30
Published:
2024-01-24
Contact:
FANG Yanming
摘要:
【目的】马尾松(Pinus massoniana)是中国亚热带地区主要的针叶树种,黄山松(P. hwangshanensis)是中国东部亚热带中山地区的主要建群种,两个物种在水平区域内分布基本重合,但垂直分布差异明显。深入了解马尾松林与黄山松林的分布格局,可以为科学管理保护马尾松和黄山松森林资源提供理论依据。【方法】基于中国亚热带东部地区8个典型马尾松林与黄山松林分布地域开展植被调查,采用非度量多维标度分析(NMDS)和多元回归树(nultivariate regression trees,MRT)探讨两种松林群落结构的差异及主要影响因子,采用线性混合效应模型定量分析各物种多样性指标对非生物因子的响应;在遵从《中国植被志》研编规范的基础上,对马尾松林与黄山松林进行初步的数量分类。【结果】马尾松群落与黄山松群落结构存在明显差异,主要影响因子为海拔;年均气温对两种群落物种丰富度影响显著,对两种群落总体物种丰富度影响最显著的因子是年均降水量,海拔对两种群落总体的均匀度产生显著的影响;对马尾松林与黄山松林进行初步植被分类,马尾松林被划分为6个群丛,黄山松林被划分为5个群丛。【结论】海拔变化塑造了中国东部地区马尾松林与黄山松林群落结构的差异,并直接或间接地影响两种群落的物种多样性。通过植被分类可以了解两种松林植被类型的多样化,并为后续两种松林分类后续工作提供参考。
中图分类号:
范明阳,胡萌,杨园,等. 中国东部地区马尾松与黄山松群落分类及群落结构和物种多样性特征[J]. 南京林业大学学报(自然科学版), 2024, 48(1): 47-58.
FAN Mingyang, HU Meng, YNAG Yuan, FANG Yanming. Community classification, structures and species diversity characteristics of Pinus massoniana and P. hwangshanensis in the eastern China[J].Journal of Nanjing Forestry University (Natural Science Edition), 2024, 48(1): 47-58.DOI: 10.12302/j.issn.1000-2006.202210029.
表1
马尾松林与黄山松林样方地理信息表"
样方 plot | 地点 site | 经度(E) / (°) longitude | 纬度(N) / (°) latitude | 海拔/m altitude | 样方plot | 地点 site | 经度(E)/ (°) longitude | 纬度(N)/ (°) latitude | 海拔/m altitude |
---|---|---|---|---|---|---|---|---|---|
P1 | 三清山 | 117.974 3 | 28.874 1 | 313 | P21 | 黄山 | 118.136 4 | 30.125 3 | 901 |
P2 | 三清山 | 117.962 3 | 28.854 5 | 498 | P22 | 黄山 | 118.137 8 | 30.125 6 | 1 000 |
P3 | 三清山 | 117.949 9 | 28.863 0 | 707 | P23 | 黄山 | 118.144 2 | 30.120 0 | 1 167 |
P4 | 三清山 | 117.975 2 | 29.881 4 | 1 086 | P24 | 黄山 | 118.146 9 | 30.132 8 | 1 720 |
P5 | 庐山 | 116.004 5 | 29.521 9 | 300 | P25 | 黄山 | 118.145 8 | 30.117 8 | 1 800 |
P6 | 庐山 | 115.983 3 | 29.537 4 | 680 | P26 | 天目山 | 119.452 8 | 30.328 3 | 450 |
P7 | 庐山 | 115.983 8 | 29.537 0 | 790 | P27 | 天目山 | 119.446 4 | 30.343 6 | 740 |
P8 | 庐山 | 115.983 9 | 29.542 9 | 900 | P28 | 天目山 | 119.450 0 | 30.338 1 | 970 |
P9 | 庐山 | 115.983 5 | 29.544 6 | 1 018 | P29 | 天目山 | 119.429 4 | 30.336 7 | 1 000 |
P10 | 庐山 | 115.973 0 | 29.572 9 | 1 155 | P30 | 天目山 | 119.453 9 | 30.329 4 | 1 000 |
P11 | 井冈山 | 114.091 5 | 26.669 8 | 480 | P31 | 天目山 | 119.430 6 | 30.349 7 | 1 300 |
P12 | 井冈山 | 114.169 2 | 26.557 3 | 730 | P32 | 凤阳山 | 119.175 2 | 27.898 8 | 1 536 |
P13 | 井冈山 | 114.948 3 | 26.344 0 | 834 | P33 | 凤阳山 | 119.179 8 | 27.891 7 | 1 660 |
P14 | 井冈山 | 114.155 4 | 26.565 2 | 920 | P34 | 凤阳山 | 119.159 9 | 27.900 7 | 1 720 |
P15 | 井冈山 | 114.108 5 | 26.545 6 | 1 112 | P35 | 四明山 | 121.123 9 | 29.777 9 | 630 |
P16 | 井冈山 | 114.121 6 | 26.601 4 | 1 300 | P36 | 四明山 | 120.989 4 | 29.634 7 | 875 |
P17 | 黄山 | 118.104 4 | 30.156 4 | 342 | P37 | 四明山 | 120.987 1 | 29.635 1 | 946 |
P18 | 黄山 | 118.110 3 | 30.158 3 | 450 | P38 | 桃花冲 | 116.033 5 | 30.979 5 | 945 |
P19 | 黄山 | 118.103 3 | 30.158 6 | 550 | P39 | 桃花冲 | 116.050 2 | 30.977 0 | 1 061 |
P20 | 黄山 | 118.105 0 | 30.158 9 | 650 | P40 | 桃花冲 | 116.059 0 | 30.977 5 | 1 160 |
表2
非生物因子对物种α多样性指数的影响"
α多样性指数 α diversity indices | 因子 factor | 马尾松群落 P. massoniana community | 黄山松群落 P. hwangshanensis community | 总体 total |
---|---|---|---|---|
物种丰富度 species richness | 海拔 | 1.12×10-5±4.75×10-4 | 1.2×10-5±4.7×10-4 | 3.4×10-4±2.3×10-4 |
年均气温 | 0.20±0.09* | 0.20±0.09* | 0.03±0.07 | |
年均降水量 | -3.68×10-4±4.75×10-4 | -3.68×10-4±8.10×10-4 | -0.002±0.001 | |
纬度 | 1.12×10-5±4.75×10-4 | 0.05±0.10 | 0.06±0.10 | |
Shannon多样性指数 Shannon diversity | 海拔 | 0.010±0.004 | 7.2×10-5±1.7×10-3 | 0.002±0.001 |
年均气温 | 0.31±0.94 | 0.87±0.57 | -0.13±0.40 | |
年均降水量 | -0.010±0.004 | 0.003±0.007 | -0.01±0.005 | |
纬度 | 0.97±0.84 | 0.44±0.93 | 0.53±0.95 | |
Gini-Simpson多样性指数 Gini-Simpson | 海拔 | 0.001±0.001 | 1.8×10-4±2.1×10-4 | 1.4×10-4±1.3×10-4* |
年均气温 | -0.04±0.10 | 0.03±0.06 | 0.01±0.04 | |
年均降水量 | -0.000 4±0.000 4 | -2.97×10-4±6.32×10-4 | -0.001 0±0.000 4 | |
纬度 | 0.04±0.09 | -0.009±0.080 | 0.02±0.06 |
表3
马尾松群系群丛分类简表"
物种 species | 群落层 community vertical layer | 特征值/% fidelity value | |||||
---|---|---|---|---|---|---|---|
PM-1(3) | PM-2(2) | PM-3(2) | PM-4(4) | PM-5(4) | PM-6(2) | ||
马银花Rhododendron ovatum | 灌木层 | 100 | 0 | 0 | 0 | 0 | 0 |
山胡椒Lindera glauca | 灌木层 | 92 | 0 | 0 | 0 | 0 | 0 |
杜鹃R. simsii | 灌木层 | 74.1 | 0 | 0 | 0 | 0 | 0 |
中国绣球Hydrangea chinensis | 灌木层 | 70.7 | 0 | 0 | 0 | 0 | 0 |
柿Diospyros kaki | 灌木层 | 70.7 | 0 | 0 | 0 | 0 | 0 |
野柿D. kaki var. silvestris | 乔木层 | 70.7 | 0 | 0 | 0 | 0 | 0 |
枫香Liquidambar formosana | 灌木层 | 70.7 | 0 | 0 | 0 | 0 | 0 |
山槐Albizia kalkora | 乔木层 | 70.7 | 0 | 0 | 0 | 0 | 0 |
赤楠Syzygium buxifolium | 灌木层 | 0 | 100 | 0 | 0 | 0 | 0 |
甜槠Castanopsis eyrei | 灌木层 | 0 | 95.9 | 0 | 0 | 0 | 0 |
木荷Schima superba | 灌木层 | 0 | 0 | 95.9 | 0 | 0 | 0 |
蕨Pteridium aquilinum var. latiusculum | 草本层 | 0 | 0 | 0 | 84.5 | 0 | 0 |
枹栎Quercus serrata | 乔木层 | 0 | 0 | 0 | 64.5 | 0 | 0 |
老鼠屎Symplocos stellaris | 乔木层 | 0 | 0 | 0 | 61.5 | 0 | 0 |
檫木Sassafras tzumu | 乔木层 | 0 | 0 | 0 | 57.7 | 0 | 0 |
狗脊Woodwardia japonica | 草本层 | 0 | 0 | 0 | 0 | 72.4 | 0 |
油茶Camellia oleifera | 灌木层 | 0 | 0 | 0 | 0 | 70.7 | 0 |
赤杨叶Alniphyllum fortunei | 乔木层 | 0 | 0 | 0 | 0 | 67.6 | 0 |
栀子Gardenia jasminoides | 灌木层 | 0 | 0 | 0 | 0 | 58.5 | 0 |
交让木Daphniphyllum macropodum | 灌木层 | 0 | 0 | 0 | 0 | 57.7 | 0 |
地棯Melastoma dodecandrum | 草本层 | 0 | 0 | 0 | 0 | 57.7 | 0 |
盐麸木Rhus chinensis | 灌木层 | 0 | 0 | 0 | 0 | 57.7 | 0 |
一年蓬Erigeron annuus | 草本层 | 0 | 0 | 0 | 0 | 0 | 100 |
南烛Vaccinium bracteatum | 灌木层 | 0 | 0 | 0 | 0 | 0 | 95.9 |
表4
黄山松群丛分类简表"
物种名 species name | 群落层 community vertical layer | 特征值/% fidelity value | |||||||
---|---|---|---|---|---|---|---|---|---|
PH-1(4) | PH-2(4) | PH-3(4) | PH-4(2) | PH-5(4) | |||||
多脉青冈Q. multinervis | 乔木层 | 72.4 | 0 | 0 | 0 | 0 | |||
小叶青冈Q. myrsinifolia | 乔木层 | 67.6 | 0 | 0 | 0 | 0 | |||
交让木D. macropodum | 灌木层 | 64.5 | 0 | 0 | 0 | 0 | |||
交让木D. macropodum | 乔木层 | 58.5 | 0 | 0 | 0 | 0 | |||
光亮山矾S. lucida | 灌木层 | 57.7 | 0 | 0 | 0 | 0 | |||
光亮山矾S. lucida | 乔木层 | 57.7 | 0 | 0 | 0 | 0 | |||
山矾S. sumuntia | 乔木层 | 57.7 | 0 | 0 | 0 | 0 | |||
马银花R. ovatum | 灌木层 | 0 | 72.9 | 0 | 0 | 0 | |||
甜槠C. eyrei | 乔木层 | 0 | 67.6 | 0 | 0 | 0 | |||
细齿叶柃Eurya nitida | 灌木层 | 0 | 63.0 | 0 | 0 | 0 | |||
鹿角杜鹃R. latoucheae | 乔木层 | 0 | 57.7 | 0 | 0 | 0 | |||
浙江红山茶C. chekiangoleosa | 灌木层 | 0 | 57.7 | 0 | 0 | 0 | |||
云锦杜鹃R. fortunei | 乔木层 | 0 | 57.7 | 0 | 0 | 0 | |||
鹿角杜鹃R. latoucheae | 灌木层 | 0 | 54.2 | 0 | 0 | 0 | |||
狭叶香港远志Polygala hongkongensis var. stenophylla | 草本层 | 0 | 0 | 57.7 | 0 | 0 | |||
丁香杜鹃R. mariesii | 灌木层 | 0 | 0 | 57.7 | 0 | 0 | |||
蛇莓Duchesnea indica | 草本层 | 0 | 0 | 57.7 | 0 | 0 | |||
紫萁Osmunda japonica | 草本层 | 0 | 0 | 0 | 100 | 0 | |||
荚蒾Viburnum dilatatum | 灌木层 | 0 | 0 | 0 | 100 | 0 | |||
山胡椒L. glauca | 灌木层 | 0 | 0 | 0 | 0 | 92.0 | |||
蕨P. aquilinum var. latiusculum | 草本层 | 0 | 0 | 0 | 0 | 92.0 | |||
灯台树Cornus controversa | 乔木层 | 0 | 0 | 0 | 0 | 58.5 | |||
山胡椒L. glauca | 乔木层 | 0 | 0 | 0 | 0 | 57.7 | |||
格药柃E. muricata | 灌木层 | 0 | 0 | 0 | 0 | 57.7 | |||
山橿L. reflexa | 灌木层 | 0 | 0 | 0 | 0 | 54.2 | |||
枹栎Q. serrata | 乔木层 | 0 | 0 | 76.3 | 0 | 76.3 | |||
枹栎Q. serrata | 灌木层 | 0 | 0 | 75.2 | 0 | 75.2 |
[1] | 王滑, 潘刚, 边巴多吉, 等. 西藏泡核桃群落结构及物种多样性分析[J]. 西部林业科学, 2015, 44(2):43-47,53. |
WANG H, PAN G, Bianbadoji, et al. The structure characteristics and species diversity of Juglans sigillata communities in Tibet Autonomous Region[J]. J West China For Sci, 2015, 44(2):43-47,53.DOI: 10.16473/j.cnki.xblykx1972.2015.02.012. | |
[2] | 柴永福, 岳明. 植物群落构建机制研究进展[J]. 生态学报, 2016, 36(15):4557-4572. |
CHAI Y F, YUE M. Research advances in plant community assembly mechanisms[J]. Acta Ecol Sin, 2016, 36(15):4557-4572. DOI: 10.5846/stxb201501140114. | |
[3] | 马克平, 黄建辉, 于顺利, 等. 北京东灵山地区植物群落多样性的研究:Ⅱ丰富度、均匀度和物种多样性指数[J]. 生态学报, 1995, 15(3):268-277. |
MA K P, HUANG J H, YU S L, et al. Plant community diversity in ongling mountain,Beijing,China:Ⅱ.species richness,evenness and species diversities[J]. Acta Ecol Sin, 1995, 15(3):268-277.DOI: 10.3321/j.issn:1000-0933.1998.01.004. | |
[4] | 郑天义, 王丹, 姬柳婷, 等. 太白山自然保护区典型森林群落数量分类、排序及多样性格局[J]. 生态学报, 2020, 40(20):7353-7361. |
ZHENG T Y, WANG D, JI L T, et al. Classification,ordination and diversity pattern of typical forest communities in Taibai Mountain Nature Reserve[J]. Acta Ecol Sin, 2020, 40(20):7353-7361.DOI: 10.5846/stxb201912252787. | |
[5] | 刘欢, 李文君, 陈杰, 等. 陕西米仓山自然保护区6种典型天然林群落的物种多样性[J]. 西北农林科技大学学报(自然科学版), 2017, 45(7):29-39. |
LIU H, LI W J, CHEN J, et al. Species diversity of six typical natural forest communities in Shaanxi Micangshan Nature Reserve[J]. J Northwest A&F Univ (Nat Sci Ed), 2017, 45(7):29-39.DOI: 10.13207/j.cnki.jnwafu.2017.07.005. | |
[6] | 陈杰, 李文君, 钟娇娇, 等. 陕西米仓山巴山冷杉天然林群落物种多样性及种群分布格局[J]. 西北农林科技大学学报(自然科学版), 2019, 47(1):69-78,89. |
CHEN J, LI W J, ZHONG J J, et al. Species diversity and spatial distribution pattern of Abies fargesii natural forest community in Shaanxi Micangshan Mountain[J]. J Northwest A&F Univ (Nat Sci Ed), 2019, 47(1):69-78,89.DOI: 10.13207/j.cnki.jnwafu.2019.01.009. | |
[7] | LEENDERTSE P C, ROOZEN A J M, ROZEMA J. Long-term changes (1953-1990) in the salt marsh vegetation at the Boschplaat on Terschelling in relation to sedimentation and flooding[J]. Plant Ecol, 1997, 132(1):49-58.DOI: 10.1023/A:1009795002076. |
[8] | BROWN R L, PEET R K. Diversity and invasibility of southern Appalachian plant communities[J]. Ecology,2003, 84(1):32-39.DOI: 10.1890/0012-9658(2003)084[0032:DAIOSA]2.0.CO;2. |
[9] | LAMB D, ERSKINE P D, PARROTTA J A. Restoration of degraded tropical forest landscapes[J]. Science, 2005, 310(5754):1628-1632.DOI: 10.1126/science.1111773. |
[10] | 张克荣, 刘应迪, 朱晓文, 等. 长沙岳麓山马尾松林的群落类型划分及物种多样性分析[J]. 林业科学, 2011, 47(4): 86-94. |
ZHANG K R, LIU Y D, ZHU X W, et al. Community types and species diversity of Pinus massoniana forests of Yuelu Mountain, Changsha[J]. Sci Silvae Sin, 2011, 47(4): 86-94. DOI: 10.11707/j.1001-7488.20110414. | |
[11] | 葛晓改, 肖文发, 曾立雄, 等. 三峡库区马尾松林土壤-凋落物层酶活性对凋落物分解的影响[J]. 生态学报, 2014, 34(9):2228-2237. |
GE X G, XIAO W F, ZENG L X, et al. Effect of soil-litter layer enzyme activities on litter decomposition in Pinus massoniana plantation in Three Gorges Reservoir Area[J]. Acta Ecol Sin, 2014, 34(9):2228-2237.DOI: 10.5846/stxb201305261185. | |
[12] | 朱政德. 中国森林的地理分布[J]. 南京林业大学学报(自然科学版), 2004, 28(1):1-8. |
ZHU Z D. The geographical distribution of forest in China[J]. J Nanjing For Univ (Nat Sci Ed), 2004, 28(1):1-8.DOI: 10.3969/j.issn.1000-2006.2004.01.001. | |
[13] | 梁赛花, 邓清华, 丁松, 等. 马尾松林林下植被多样性、生物量研究进展[J]. 南方林业科学, 2015, 43(1):30-31,46. |
LIANG S H, DENG Q H, DING S, et al. A review on diversity and biomass of understory vegetation in Pinus massoniana forest[J]. Nanfang For Sci, 2015, 43(1):30-31,46.DOI: 10.16259/j.cnki.36-1342/s.2015.01.009. | |
[14] | 刘艳会, 刘金福, 何中声, 等. 基于戴云山固定样地黄山松群落物种组成与结构研究[J]. 广西植物, 2017, 37(7):881-890. |
LIU Y H, LIU J F, HE Z S, et al. Pinus taiwanensis community composition and structure based on fixed sample Daiyun Mountain[J]. Guihaia, 2017, 37(7):881-890.DOI: 10.11931/guihaia.gxzw201607015. | |
[15] | 苏松锦, 刘金福, 兰思仁, 等. 黄山松研究综述(1960—2014)及其知识图谱分析[J]. 福建农林大学学报(自然科学版), 2015, 44(5):478-486. |
SU S J, LIU J F, LAN S R, et al. A review of Pinus taiwanensis studies(1960-2014) and the knowledge domain analysis[J]. J Fujian Agric For Univ (Nat Sci Ed), 2015, 44(5):478-486.DOI: 10.13323/j.cnki.j.fafu(nat.sci.).2015.05.006. | |
[16] | GERNANDT D S, LOPEZ G G, GARCIA S O, et al. Phylogeny and classification of Pinus[J]. Taxon, 2005, 54(1): 29-42. DOI: 10.2307/25065300. |
[17] | 童再康, 范义荣. 黄山松分布区气候生态区划[J]. 中南林学院学报, 1993(1):81-87. |
TONG Z K, FAN Y R. Dividing the distribution area of Pinus taiwanensis based on climatic and ecologic factors[J]. J Central South For Univ, 1993(1):81-87.DOI:10.14067/j.cnki.1673-923x.1993.01.016. | |
[18] | 马元屾, 王中生, 余华, 等. 不同海拔梯度下黄山松与马尾松针叶形态·光合生理特性的研究[J]. 安徽农业科学, 2012, 40(29):14155-14158,14173. |
MA Y S, WANG Z S, YU H, et al. A study in photosynthetic physiological and morphologic parameters of needles of Pinus taiwanensis and Pinus massoniana in different elevation along altitudinal gradients[J]. J Anhui Agric Sci, 2012, 40(29):14155-14158,14173.DOI: 10.13989/j.cnki.0517-6611.2012.29.305. | |
[19] | 唐志尧, 方精云. 植物物种多样性的垂直分布格局[J]. 生物多样性, 2004, 12(1):20-28. |
TANG Z Y, FANG J Y. A review on the elevational patterns of plant species diversity[J]. Chin Biodivers, 2004, 12(1):20-28.DOI: 10.3321/j.issn:1005-0094.2004.01.004. | |
[20] | FONDA R W, BLISS L C. Forest vegetation of the Montane and Subalpine Zones, Olympic Mountains, Washington[J]. Ecological Monographs, 1969, 39(3): 271-301.DOI: 10.2307/1948547. |
[21] | WHITTAKER R J, WILLIS K J, FIELD R. Scale and species richness:towards a general,hierarchical theory of species diversity[Review][J]. J Biogeogr, 2001, 28(4):453-470.DOI: 10.1046/j.1365-2699.2001.00563.x. |
[22] | EMERSON B C, GILLESPIE R G. Phylogenetic analysis of community assembly and structure over space and time[J]. Trends Ecol Evol, 2008, 23(11):619-630.DOI: 10.1016/j.tree.2008.07.005. |
[23] | CURRIE D J, MITTELBACH G G, CORNELL H V, et al. Predictions and tests of climate-based hypotheses of broad-scale variation in taxonomic richness[J]. Ecol Lett, 2004, 7(12):1121-1134.DOI: 10.1111/j.1461-0248.2004.00671.x. |
[24] | RAHBEK C. The role of spatial scale and the perception of large-scale species-richness patterns[J]. Ecol Lett, 2005, 8(2):224-239.DOI: 10.1111/j.1461-0248.2004.00701.x. |
[25] | 吴征镒. 中国植被[M]. 北京: 科学出版社, 1980: 217-230. |
[26] | 吴中伦. 安徽黄山黄山松的初步观察[J]. 林业科学, 1963, 8 (2): 114-126. |
WU Z L. Note on Hwangshan pine of Hwangshan, Anhwei Province[J]. Scientia Silvae Sinicae, 1963, 8 (2): 114-126. | |
[27] | 罗世家, 刘永清. 黄山松群丛特征的比较[J]. 湖北民族学院学报(自然科学版), 1999(3): 40-43. |
LUO S J, LIU Y Q. Characteristic comparision of Huangshan pine (Pinus hwangshanensis) community[J]. Journal of Hubei Institute for Nationalities (Natural Science Edition), 1999(3): 40-43. DOI: CNKI:SUN:HBXZ.0.1999-03-010. | |
[28] | 林鹏, 叶庆华. 武夷山植被研究(四) 黄岗山针叶林[J]. 武夷科学, 1984(1):23-30. |
LIN P, YE Q H. The subtropic coniferous vegetation of Huang-Gang Mountain in Fujian,China[J]. Wuyi Sci J, 1984(1):23-30. | |
[29] | JIANG L, HE Z S, GU X G, et al. Classification and ordination of the Pinus taiwanensis forest on Daiyun Mountain, Fujian Province, China[J]. Taiwania, 2020, 65(2): 119-128. DOI:10.6165/tai.2020.65.119. |
[30] | 余小平, 李新. 重庆南山马尾松群落的数量分类研究[J]. 重庆师范学院学报(自然科学版), 1992(1): 55-60,81. |
YU X P, LI X. Quantitative classification of masson pine community at Nan Mountain in Chongqing[J]. Journal of Chongqing teachers college (Natural Science Edition), 1992(1): 55-60,81. | |
[31] | 方炜, 彭少麟. 鼎湖山马尾松群落演替过程物种变化之研究[J]. 热带亚热带植物学报, 1995, 3(4):30-37. |
FANG W, PENG S L. Changes of tree species in the succession process of Pinus massoniana community in Dinghushan,Guangdong,P.R.China[J]. J Trop Subtrop Bot, 1995, 3(4):30-37.DOI: CNKI:SUN:RYZB.0.1995-04-004. | |
[32] | 祁丽霞, 刘金福, 黄嘉航, 等. 戴云山黄山松、马尾松针叶抗氧化酶活性的海拔梯度分布格局[J]. 福建农林大学学报(自然科学版), 2016, 45(1):35-41. |
QI L X, LIU J F, HUANG J H, et al. The altitudinal gradient pattern of antioxidant enzyme activity in Pinus taiwanensis and Pinus massoniana needles[J]. J Fujian Agric For Univ (Nat Sci Ed), 2016, 45(1):35-41. DOI: 10.13323/j.cnki.j.fafu(Nat.Sci.).2016.01.007. | |
[33] | 罗世家, 邹惠渝, 梁师文. 黄山松与马尾松基因渐渗的研究[J]. 林业科学, 2001, 37(6): 118-122. |
LUO S J, ZOU H Y, LIANG S W. Study on the introgressive hybridization between Pinus hwangshanensis and P.massoniana[J]. Sci Silvae Sin, 2001, 37(6): 118-122. DOI: 10.11707/j.1001-7488.20010622. | |
[34] | 翟大才, 宣磊, 周琦, 等. 黄山地区马尾松和黄山松基于SSR标记的基因渐渗研究[J]. 分子植物育种, 2018, 16(14):4614-4622. |
ZHAI D C, XUAN L, ZHOU Q, et al. Genetic introgression of Pinus massoniana and Pinus taiwanensis in Huangshan area based on SSR markers[J]. Mol Plant Breed, 2018, 16(14):4614-4622.DOI: 10.13271/j.mpb.016.004614. | |
[35] | 张利锐. 马尾松和黄山松群体遗传学与物种分化研究[D]. 兰州: 兰州大学, 2011. |
ZHANG L R. Study on population genetics and species differentiation of Pinus massoniana and Pinus taiwanensis[D]. Lanzhou: Lanzhou University, 2011. | |
[36] | 王国宏, 方精云, 郭柯, 等. 《中国植被志》研编内容与规范[J]. 植物生态学报, 2020, 44(2):128-178. |
WANG G H, FANG J Y, GUO K, et al. Contents and protocols for the classification and description of vegetation formations,alliances and associations of vegetation of China[J]. Chin J Plant Ecol, 2020, 44(2):128-178.DOI: 10.17521/cjpe.2019.0272. | |
[37] | 方精云, 王襄平, 沈泽昊, 等. 植物群落清查的主要内容、方法和技术规范[J]. 生物多样性, 2009, 17(6):533-548. |
FANG J Y, WANG X P, SHEN Z H, et al. Methods and protocols for plant community inventory[J]. Biodivers Sci, 2009, 17(6):533-548. DOI:10.3724/SP.J.1003.2009.09253. | |
[38] | CHAO A. Estimating the population size for capture-recapture data with unequal catchability[J]. Biometrics, 1987, 43(4):783-791. |
[39] | GASTON K J. Global patterns in biodiversity[J]. Nature, 2000, 405(6783):220-227.DOI: 10.1038/35012228. |
[40] | ABBAS S, NICHOL J E, ZHANG J L, et al. The accumulation of species and recovery of species composition along a 70-year succession in a tropical secondary forest[J]. Ecol Indic, 2019, 106:105524.DOI: 10.1016/j.ecolind.2019.105524. |
[41] | 解雷, 陈浩, 赵荣, 等. 黄山常绿阔叶林群丛数量分类及物种多样性格局——以10.24 hm2森林动态监测样地为例[J]. 陕西师范大学学报(自然科学版), 2023, 51(3):86-95. |
XIE L, CHEN H, ZHAO R, et al. Association classification and species diversity of the subtropical evergreen broad-leaved forest in Huangshan, Anhui, China: a case study of 10.24 hm2 forest dynamics plot[J]. J Shannxi Norm Univ (Nat Sci Edi), 2023, 51(3):86-95. DOI: 10.15983/j.cnki.jsnu.2023207. | |
[42] | 马克平, 叶万辉, 于顺利, 等. 北京东灵山地区植物群落多样性研究Ⅷ:群落组成随海拔梯度的变化[J]. 生态学报, 1997, 17(6):593-600. |
MA K P, YE W H, YU S L, et al. Studies on plant community diversity in Dongling Mountain,Beijing,China Ⅷ: variation of composition along elevational gradient[J]. Acta Ecol Sin, 1997, 17(6):593-600. | |
[43] | WANG Z H, RAHBEK C, FANG J Y. Effects of geographical extent on the determinants of woody plant diversity[J]. Ecography, 2012, 35(12):1160-1167.DOI: 10.1111/j.1600-0587.2012.07786.x. |
[44] | 方文静, 蔡琼, 朱江玲, 等. 华北地区落叶松林的分布、群落结构和物种多样性[J]. 植物生态学报, 2019, 43(9):742-752. |
FANG W J, CAI Q, ZHU J L, et al. Distribution,community structures and species diversity of larch forests in north China[J]. Chin J Plant Ecol, 2019, 43(9):742-752.DOI: 10.17521/cjpe.2018.0244. | |
[45] | 吴昊. 秦岭松栎林群落物种丰富度特征及其环境解释[J]. 生态环境学报, 2017, 26(6):931-938. |
WU H. Species richness and environmental interpretation of pine and oak forest community in Qinling Mountains[J]. Ecol Environ Sci, 2017, 26(6):931-938.DOI: 10.16258/j.cnki.1674-5906.2017.06.004. | |
[46] | SCHERRER D, KÖRNER C. Topographically controlled thermal-habitat differentiation buffers alpine plant diversity against climate warming[J]. J Biogeogr, 2011, 38(2):406-416.DOI: 10.1111/j.1365-2699.2010.02407.x. |
[47] | ZHANG J L, ZHOU Y, ZHOU G S, et al. Structure and composition of natural Gmelin larch (Larix gmelinii var. gmelinii) forests in response to spatial climatic changes[J]. PLoS One, 2013, 8(6):e66668.DOI: 10.1371/journal.pone.0066668. |
[48] | TAO S L, GUO Q H, LI C, et al. Global patterns and determinants of forest canopy height[J]. Ecology, 2016, 97(12):3265-3270.DOI: 10.1002/ecy.1580. |
[49] | 刘开明, 郑智, 龚大洁. 物种丰富度的垂直分布格局及其形成机制[J]. 生态学杂志, 2017, 36(2):541-554. |
LIU K M, ZHENG Z, GONG D J. Elevational patterns of species richness and their underlying mechanism[J]. Chin J Ecol, 2017, 36(2):541-554.DOI: 10.13292/j.1000-4890.201702.015. | |
[50] | 朱德煌. 戴云山黄山松种群特征及其环境因子研究[D]. 福州: 福建农林大学, 2013. |
ZHU D H. Study on population characteristics and environmental factors of Pinus taiwanensis in Daiyun Mountain[D]. Fuzhou: Fujian Agriculture and Forestry University, 2013. | |
[51] | 李贺, 张维康, 王国宏. 中国云杉林的地理分布与气候因子间的关系[J]. 植物生态学报, 2012, 36(5):372-381. |
LI H, ZHANG W K, WANG G H. Relationship between climatic factors and geographical distribution of spruce forests in China[J]. Chin J Plant Ecol, 2012, 36(5):372-381.DOI: 10.3724/SP.J.1258.2012.00372. | |
[52] | 孟苗婧, 张金池, 郭晓平, 等. 海拔变化对黄山松阔叶混交林土壤有机碳组分的影响[J]. 南京林业大学学报(自然科学版), 2018, 42(6):106-112. |
MENG M J, ZHANG J C, GUO X P, et al. Effects of altitude change on soil organic carbon fractions in Pinus taiwanensis and broad-leaved mixed forest[J]. J Nanjing For Univ (Nat Sci Ed), 2018, 42(6):106-112.DOI: 10.3969/j.issn.1000-2006.201712031. | |
[53] | 祁丽霞. 戴云山黄山松-马尾松叶性状对海拔梯度的响应研究[D]. 福州: 福建农林大学, 2015. |
QI L X. Response of leaf characters of Pinus taiwanensis-Pinus massoniana in Daiyun Mountain to altitude gradient[D]. Fuzhou: Fujian Agriculture and Forestry University, 2015. | |
[54] | 李淑娴. 马尾松和黄山松物种分化遗传机制研究[D]. 南京: 南京林业大学, 2012. |
LI S X. Study on genetic mechanism of species differentiation of Pinus massoniana and Pinus taiwanensis[D]. Nanjing: Nanjing Forestry University, 2012. | |
[55] | 任国学, 刘金福, 徐道炜, 等. 戴云山国家级自然保护区黄山松群落类型与物种多样性分析[J]. 植物资源与环境学报, 2011, 20(3):82-88. |
REN G X, LIU J F, XU D W, et al. Analysis on classification and species diversity of Pinus taiwanensis community in Daiyun Mountain National Nature Reserve[J]. J Plant Resour Environ, 2011, 20(3):82-88. DOI: 10.3969/j.issn.1674-7895.2011.03.013. | |
[56] | 刘金福, 朱德煌, 兰思仁, 等. 戴云山黄山松群落与环境的关联[J]. 生态学报, 2013, 33(18):5731-5736. |
LIU J F, ZHU D H, LAN S R, et al. Association between environment and community of Pinus taiwanensis in Daiyun Mountain[J]. Acta Ecol Sin, 2013, 33(18):5731-5736.DOI: 10.5846/stxb201304270832. |
[1] | 颜铮明, 阮宏华, 廖家辉, 石珂, 倪娟平, 曹国华, 沈彩芹, 丁学农, 赵小龙, 庄鑫. 不同林龄杨树人工林地表甲虫群落多样性特征[J]. 南京林业大学学报(自然科学版), 2023, 47(6): 236-242. |
[2] | 尹世轩, 吴永波, 黄潇宇. 常州市金坛生态红线区鸟类多样性研究[J]. 南京林业大学学报(自然科学版), 2023, 47(4): 219-225. |
[3] | 郭丽丽, 张晨洁, 王菲, 沈佳佳, 张凯月, 何丽霞, 郭琪, 侯小改. 牡丹野生种根际土壤细菌群落特征分析[J]. 南京林业大学学报(自然科学版), 2023, 47(3): 45-55. |
[4] | 伊贤贵, 董鹏, 谢春平, 彭智奇, 杨国栋, 董京京, 钟育谦, 翟飞飞, 王贤荣. 江苏宜兴龙池山自然保护区固定样地物种组成分析[J]. 南京林业大学学报(自然科学版), 2021, 45(6): 159-168. |
[5] | 王玄, 崔鹏, 丁晶晶, 常青. 江苏南部沿海越冬水鸟群落结构及多样性分析[J]. 南京林业大学学报(自然科学版), 2021, 45(5): 178-184. |
[6] | 陈宏健, 郝德君, 田敏, 周杨, 夏小洪, 赵欣怡, 乔恒, 谈家金. 室内饲养松墨天牛幼虫不同肠段细菌的群落结构及功能分析[J]. 南京林业大学学报(自然科学版), 2021, 45(3): 143-151. |
[7] | 朱晗, 郝德君, 魏原芝, 孙丽昕, 文全民. 仁扇舟蛾幼虫肠道可培养细菌群落结构分析[J]. 南京林业大学学报(自然科学版), 2021, 45(2): 171-176. |
[8] | 陈秀波, 段文标, 陈立新, 朱德全, 赵晨晨, 刘东旭. 小兴安岭3种原始红松混交林土壤nirK型反硝化微生物群落特征[J]. 南京林业大学学报(自然科学版), 2021, 45(2): 77-86. |
[9] | 周永晟, 徐子恒, 袁发银, 尚旭岚, 孙操稳, 方升佐. 亚热带3个地点青钱柳群落特征比较[J]. 南京林业大学学报(自然科学版), 2021, 45(1): 29-35. |
[10] | 潘婷婷, 陈林, 杨国栋, 伊贤贵, 王贤荣. 南京北部郊野森林群落物种多样性及其环境解释[J]. 南京林业大学学报(自然科学版), 2020, 44(6): 48-54. |
[11] | 罗碧珍, 胡海清, 罗斯生, 魏书精, 吴泽鹏, 刘菲. 林火干扰对广东马尾松林土壤有机碳密度及其活性有机碳的影响[J]. 南京林业大学学报(自然科学版), 2020, 44(5): 132-140. |
[12] | 吴思思, 鲁小珍, 王馨悦, 葛之葳. 两种类型人为干扰对城市风景林植物多样性的影响[J]. 南京林业大学学报(自然科学版), 2019, 43(5): 128-134. |
[13] | 伊贤贵,丁晖,方炎明,叶要清,陈水飞,王旭,从睿,张开文,李垚,王贤荣. 基于固定样地的黄山不同海拔森林群落物种多样性分析[J]. 南京林业大学学报(自然科学版), 2018, 42(01): 149-155. |
[14] | 杨宝玲,张文文,范换,王邵军,阮宏华,沈彩芹,曹国华. 苏北沿海地区不同土地利用类型下土壤动物群落结构特征[J]. 南京林业大学学报(自然科学版), 2017, 41(06): 120-126. |
[15] | 周之栋,卜晓莉,吴永波,薛建辉. 生物炭对土壤微生物特性影响的研究进展[J]. 南京林业大学学报(自然科学版), 2016, 40(06): 1-8. |
阅读次数 | ||||||
全文 |
|
|||||
摘要 |
|
|||||