凋落物层是森林土壤不可缺少的保护层,在森林水土保持及水源涵养中起着不可替代的作用。为探究中国东北东部山地不同演替阶段针阔混交林凋落物持水特性的变化规律,以及持水特性与演替阶段之间的关系,以吉林省蛟河市林业实验区管理局林场不同演替阶段针阔混交林凋落物为研究对象,分别在中龄林、近熟林和成熟林固定监测样地中采用蛇形取样法均匀地选择5个边长为20 cm的正方形样方,并将每个样方内的未分解层和半分解层的凋落物取回实验室进行持水特性分析。采用室内浸水法计算凋落物的蓄积量、持水量、持水率以及凋落物的有效拦蓄量,并结合野外观测数据分析林下凋落物的持水特性与演替阶段之间的关系。结果表明:不同演替阶段凋落物的总蓄积量为成熟林(7.26 t/hm2)>近熟林(4.56 t/hm2)>中龄林(3.68 t/hm2),持水量大小为成熟林(21.23 t/hm2)<近熟林(35.24 t/hm2)<中龄林(47.71 t/hm2),持水率大小为成熟林(844.72%)>近熟林(742.58%)>中龄林(592.02%),有效拦蓄量为成熟林(31.32 t/hm2)>近熟林(20.52 t/hm2)>中龄林(11.98 t/hm2)。林分各演替阶段不同分解程度的凋落物持水量与持水率均随着浸水时间的增加呈对数关系增长,吸水速率则随浸水时间的增加呈幂指数关系下降。研究表明,成熟林的持水性能最强,近熟林、中龄林持水性能依次减弱。不同演替阶段林分最大持水量、最大持水率与蓄积量大小排序一致,表明持水量、持水率与蓄积量有很强的相关关系,蓄积量越大,凋落物持水量、持水率越高,森林凋落物持水能力越强。
Abstract
The forest litter layer is indispensible for the protection of forest soil, it plays an important role in forest soil and water conservation. The purpose of this paper is to analyze the variation laws of hydrological characteristics of forest litters in conifer and broad-leaved mixed forests at different successional stages in the northeast mountainous area of China. We investigated the forest litters in conifer and broad-leaved mixed forests at different successional stages in Jiaohe management bureau of forestry experimental area, Jilin Province. The litter samples were collected from middle-aged forest, near-mature forest and mature forest. Within each forest plot, a total of five square subplots with dimension of 20 cm×20 cm were set up using serpentine sampling method. The litters in the un-decomposed layer and semi-decomposed layer were taken back to lab for analyzing the hydrological characteristics of forest litters. The standard branch soaking method was used to calculate the storage of forest litters, water holding capacity, water holding rate, as well as the modified interception amount. For each sample, the litters were divided into four groups evenly and part of the litters was selected to measure the weight by electronic analytical balance, and then put in the 100-mesh nylon mesh. The soaking time was set as 0.25, 0.5, 1, 2, 4, 8 and 24 h, respectively. Combined with the field observation data, the relationship between the hydrological characteristics of forest litters and successional stages was analyzed. The litter storage of mature forest was the largest(7.26 t/hm2), followed by near-mature forest(4.56 t/hm2), and middle age forest(3.68 t/hm2).The order of the maximum water holding capacity was as follows: mature forest(21.23 t/hm2)>near-mature forest(35.24 t/hm2)>middle-aged forest(47.71 t/hm2)and the maximum water holding rate was 844.72% in mature forest, 742.58% in near-mature forest, and 592.02% in middle-aged forest, respectively.The descending order of the modified interception amount was mature forest(31.32 t/hm2), near-mature forest(20.52 t/hm2), and middle-aged forest(11.98 t/hm2). The soaking experiment demonstrated that both water holding capacity and water holding rate had remarkable logarithmic correlation with soaking time, while the absorption rate and soaking time was of a power function. The water holding capacity of mature forest was the largest, followed by near-mature forest and middle age forest. This is due to the fact that along with the forest succession the forest crown tends to close, the understory species becomes rich and nutrients increase, therefore the water holding capacity of forest litter layers increase gradually. The orders of maximum water holding capacity, maximum water holding rate and litter storage volume were consistent among different forest successional stages, which indicated that the there were strong correlation between water holding capacity and water holding rate with litter storage volume. The higher the litter storage volume, the higher the water holding capacity and water holding rate.
{{custom_sec.title}}
{{custom_sec.title}}
{{custom_sec.content}}
参考文献
[1] Liu C J, Iivesniemi H, Berg B, et al. Aboveground litterfall in Eurasian forests[J]. Journal of Forestry Research, 2003, 14(1): 27-34.
[2] 赵晓春, 刘建军, 任军辉, 等. 贺兰山4种典型森林类型凋落物持水性能研究[J]. 水土保持研究, 2011, 18(2):107-111.
Zhao X C, Liu J J, Ren J H, et al. Study on water-holding capacity of litter in four forest types of Helan Mountain[J].Research of Soil and Water Conservation, 2011, 18(2): 107-111.
[3] 郭辉, 董希斌, 蒙宽宏, 等. 小兴安岭低质林采伐改造后枯落物持水特性变化分析[J]. 林业科学, 2010, 46(6):146-153.
Guo H, Dong X B, Meng K H, et al. Analysis about change of water-holding characteristic of litter layer after logging reform in low-quality forest stands of Lesser Khingan Range[J]. Scientia Silvae Sinicae, 2010, 46(6):146-153.
[4] 周娟, 陈丽华, 郭文体, 等. 大辽河流域水源涵养林枯落物持水特性研究[J]. 水土保持通报, 2013, 33(4):136-141.
Zhou J, Chen L H, Guo W T,et al. Water-holding capacity of litter of forest for water conservation in Great Liaohe River basin[J]. Bulletin of Soil and Water Conservation, 2013, 33(4):136-141.
[5] 李超, 董希斌, 宋启亮. 大兴安岭白桦低质林皆伐改造后枯落物水文效应[J]. 东北林业大学学报, 2013, 41(10):23-27.
Li C, Dong X B, Song Q L. Hydrological effect of birch coppice low-quality forest litter after clear-cutting transformation in Daxing'an Mountains[J]. Journal of Northeast Forestry University, 2013, 41(10):23-27.
[6] Waring R H, Schlesinger W H. Forest ecosystems. concepts and management[M]. New York: Academic Press, 1985:95-101.
[7] Jonczak J. Dynamics, structure and properties of plant litterfall in a 120-year old beech stand in Middle Pomerania between 2007-2010[J]. Soil Sci Annu, 2013, 64(1):8-13.
[8] Arunachalam A, Arunachalam K M, Pandey H N, et al. Fine litterfall and nutrient dynamics during forest regrowth in the humid subtropics of north-eastern India[J]. Forest Ecology and Management, 1998, 110(1): 209-219.
[9] 魏强, 张广忠, 凌雷, 等. 甘肃兴隆山主要森林类型凋落物及土壤层的蓄水功能[J]. 南京林业大学学报(自然科学版), 2013, 37(2):78-84.Doi:10.3969/j.issn.1000-2006.2013.02.014.
Wei Q, Zhang G Z, Ling L, et al. Water conservation function of litter and soil layer under main forest types in Xinglong Mountain of Gansu[J]. Journal of Nanjing Forestry University(Natural Sciences Edition), 2013, 37(2):78-84.
[10] 常雅军, 曹靖, 马建伟, 等. 秦岭西部山地针叶林凋落物持水特性[J]. 应用生态学报, 2008, 19(11):2346-2351.
Chang Y J,Cao J, Ma J W, et al. Water-holding characteristics of coniferous forest litters in west Qinling Mountains[J]. Chinese Journal of Applied Ecology, 2008, 19(11):2346-2351.
[11] Guevara-Escobar A, Gonzalez-Sosa E, Ramos-Salinas M, et al. Experimental analysis of drainage and water storage of litter layers[J]. Hydrol Earth Syst Sci, 2007, 11(5):1703-1716.
[12] Marin C T, Bouten I W, Dekker S. Forest floor water dynamics and root water uptake in four forest ecosystems in northwest Amazonia[J]. J Hydrolo, 2000, 237(3):169-183.
[13] Vitale M, Savi F, Baldantoni D, et al. Modeling of early stage litter decomposition in Mediterranean mixed forests: functional aspects affected by local climate[J]. Journal of Biogeosciences and Forestry, 2014, 8:517-525.
[14] 任向荣, 薛立, 曹鹤, 等. 3种人工林凋落物的持水特性[J]. 华南农业大学学报, 2008, 29(3):47-51.
Ren X R, Xue L,Cao H, et al. Water-holding characteristics of litter in three plantations[J]. Journal of South China Agriculture University, 2008, 29(3):47-51.
[15] 樊登星, 余新晓, 岳永杰, 等. 北京西山不同林分枯落物层持水特性研究[J]. 北京林业大学学报, 2008, 30(S2):177-181.
Fan D X, Yu X X, Yue Y J,et al. Water-holding characteristics of litter layer of different forest stands in Xishan Mountain in Beijing[J]. Journal of Beijing Forestry University, 2008, 30(S2):177-181.
[16] 卢洪健, 李金涛, 刘文杰. 西双版纳橡胶林枯落物的持水性能与截留特征[J]. 南京林业大学学报(自然科学版), 2011, 35(4):67-73.Doi:10.3969/j.issn.1000-2006.2011.04.014.
Lu H J, Li J T, Liu W J. Study on water-holding capability and interception characteristics of litter layers under rubber plantations in Xishuangbanna, southwestern China[J]. Journal of Nanjing Forestry University(Natural Sciences Edition), 2011, 35(4):67-73.
[17] 郭汉清, 韩有志, 白秀梅. 不同林分枯落物水文效应和地表糙率系数研究[J]. 水土保持学报,2010, 24(2):179-183.
Guo H Q, Han Y Z, Bai X M. Hydrological effects of litter on different forest stands and study about surface roughness coefficient[J]. Journal of Soil and Water Conservation, 2010, 24(2):179-183.
[18] 顾宇书, 邢兆凯, 韩友志, 等. 浑河上游4种典型林分类型枯落物持水特征[J]. 南京林业大学学报(自然科学版), 2013, 37(1):31-36.Doi:10.3969/j.issn.1000-2006.2013.01.005.
Gu Y S, Xing Z K, Han Y Z, et al. Water-holding characteristics of four typical forests litter in upstream of Hunhe river[J]. Journal of Nanjing Forestry University(Natural Sciences Edition), 2013, 37(1):31-36.
[19] 姜俊, 张春雨, 赵秀海. 吉林蛟河42hm2针阔混交林样地植物种面积关系[J]. 植物生态学报,2012, 36(1):30-38.
Jiang J, Zhang C Y, Zhao X H. Plant species-area relationship in a 42-hm2 research plot of coniferous and board-leaved mixed forest in Jiaohe, Jilin Province, China[J]. Chinese Journal of Plant Ecology, 2012, 36(1):30-38.
[20] 何怀江, 张春雨, 赵秀海, 等. 吉林蛟河次生针阔混交林群落结构与微地形关系[J]. 东北林业大学学报,2014,42(5):13-17,33.
He H J, Zhang C Y, Zhao X H, et al. Associations between community structure and microtopogrphy in mixed broadleaf-conifer forest of Jiaohe, Jilin Province[J]. Journal of Northeast Forestry University, 2014, 42(5):13-17,33.
[21] 吴金卓, 蔡小溪, 林文树. 吉林蛟河不同演替阶段针阔混交林土壤健康评价[J]. 东北林业大学学报,2015,43(6):78-82.
Wu J Z, Cai X X, Lin W S. Assessment on the soil health of conifer and broad-leaved mixed forests at different successional stages in Jiaohe, Jilin Province [J]. Journal of Northeast Forestry University,2015, 43(6):78-82.
[22] 代海军. 吉林蛟河阔叶红松林主要树种异速生长模型[D]. 北京: 北京林业大学, 2013.
Dai H J. Allometric models of dominant tree species in korean pine broadleaf forest in Jiaohe, Jilin Province[D]. Beijing: Beijing Forestry University, 2013.
[23] 丁胜建. 老龄阔叶红松林主要树种空间分布及生境关联分析[D]. 北京: 北京林业大学, 2012:14.
Ding S J. Habitat associations of dominant species spatial distribution in old growth broad-leaved Korean pine(Pinus koraiensis)forest[D]. Beijing: Beijing Forestry University, 2012:14.
[24] Anderson-Teixeira K J,Davies S J,Bennett A C,et al. CTFS-Forest GEO:a worldwide network monitoring forests in an era of global change[J]. Global Change Biology, 2015, 21: 528-549.
[25] 彭萱亦. 不同演替阶段针阔混交林生物多样性评价指标体系研究[D]. 哈尔滨:东北林业大学, 2014:7-9.
Peng X Y. Study on forest biodiversity evaluation system for conifer and broad-leaved mixed forest at different successional stages [D]. Harbin: Northeast Forestry University, 2014:7-9.
[26] 刘尚华, 冯朝阳, 吕世海, 等. 京西百花山区6种植物群落凋落物持水性能研究[J]. 水土保持学报, 2007, 21(6):179-182.
Liu S H, Feng Z Y, LV S H,et al. Study on water-holding capacity of litter in six vegetation communities in Baihua Mountains Areas of West Beijing[J]. Journal of Soil and Water Conservation, 2007, 21(6):179-182.
[27] 贺宇, 丁国栋, 臧荫桐, 等. 燕山山地典型森林枯落物持水特性[J]. 四川农业大学学报, 2012, 30(2):161-166.
He Y, Ding G D, Zang Y T, et al. Study on water-holding characteristics of typical forest litter in Yanshan Mountain[J]. Journal of Sichuan Agricultural University, 2012, 30(2):161-166.
[28] 田野宏,满秀玲,李奕,等. 大兴安岭北部天然次生林枯落物及土壤水文功能研究[J]. 水土保持学报, 2013, 27(6):113-118+129.
Tian Y H, Man X L, Li Y, et al. Research on litter and soil hydrology functions of natural secondary forests in North of Da Xing'an Mountains[J]. Journal of Soil and Water Conservation, 2013, 27(6):113-118,129.
[29] 单洪伟,李文影. 白桦次生林4个林龄0~30 cm土层水源涵养功能比较[J]. 森林工程, 2014, 30(1):41-45.
Shan H W, Li W Y. Comparison of water conservation function in 0-30cm soil layer of Betula platyphylla secondary forest at four stages[J]. Forest Engineering, 2014, 30(1):41-45.
[30] 杨玉莲, 马兴艳, 吴庆贵, 等. 龙门山断裂带主要森林类型凋落物累积量及其持水特性[J]. 水土保持通报, 2014, 34(3): 69-74.
Yang Y L, Ma X Y, Wu Q G, et al. Accumulation amount of litters and their water-holding characteristics under main forest types in Longmen Mountains fault zone[J]. Bulletin of Soil and Water Conservation, 2014, 34(3): 69-74.
[31] 刘效东,乔玉娜,周国逸,等. 鼎湖山3种不同演替阶段森林凋落物的持水特性[J]. 林业科学,2013,49(9):8-15.
Liu X D, Qiao Y N, Zhou G Y, et al.Water-holding characteristics of litters in three forests at different successional stages in Dinghushan[J]. Scientia Silvae Sinicae, 2013,49(9):8-15.
[32] 雷云飞,张卓文,苏开君,等. 流溪河森林各演替阶段凋落物层的水文特性[J]. 中南林业科技大学学报, 2007, 27(6):38-43.
Lei Y F, Zhang Z W, Su K J, et al. Hydrological characteristics of the litter form in different forest succession periods at Liuxihe watershed[J]. Journal of Central South University of Forestry & Technology, 2007, 27(6):38-43.
基金
收稿日期:2015-04-29 修回日期:2015-08-28
基金项目:中央高校基本科研业务费专项资金项目(2572015CA01); 国家自然科学基金项目(31400539); “十二五”国家科技支撑计划(2012BAC01B03)
第一作者:吴金卓(wujinzhuo1980@163.com),副教授。*通信作者:林文树(linwenshu@126.com),副教授。
引文格式:吴金卓,孔琳琳,王娇娇,等. 吉林蛟河不同演替阶段针阔混交林凋落物持水特性研究[J]. 南京林业大学学报(自然科学版),2016,40(2):113-120.