JOURNAL OF NANJING FORESTRY UNIVERSITY ›› 2021, Vol. 45 ›› Issue (1): 189-196.doi: 10.12302/j.issn.1000-2006.202005008
Previous Articles Next Articles
WANG Lujun1(), CAI Chunju1,2,*(
), TANG Xiaolu3, FAN Shaohui1
Received:
2020-05-07
Accepted:
2020-06-25
Online:
2021-01-30
Published:
2021-02-01
Contact:
CAI Chunju
E-mail:wuatang@icbr.ac.cn;caicj@icbr.ac.cn
CLC Number:
WANG Lujun, CAI Chunju, TANG Xiaolu, FAN Shaohui. Aboveground biomass allocation patterns and model construction of Bambusa rigida[J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2021, 45(1): 189-196.
Table 1
Site conditions and stand status"
编号 No. | 海拔/m altitude | 坡度/(°) slope | 坡向 aspect | 丛密度/ (丛·hm-2) clump density | 丛株数 quantity | 平均株高/m mean plant height | 平均胸径/cm mean DBH |
---|---|---|---|---|---|---|---|
1 | 300 | 30 | 阳坡 | 907 | 7.6 | 12.03±1.89 | 4.83±0.73 |
2 | 312 | 35 | 阳坡 | 852 | 8.3 | 11.78±2.01 | 4.89±0.79 |
3 | 333 | 33 | 阳坡 | 1 500 | 7.0 | 12.35±2.45 | 4.85±0.75 |
4 | 290 | 28 | 阳坡 | 1 425 | 7.2 | 12.30±2.35 | 4.85±0.73 |
5 | 325 | 29 | 阳坡 | 852 | 7.5 | 12.55±2.20 | 4.90±0.83 |
Table 2
11 kinds of commonly used biomass models"
模型类型 model type | 自变量 independent variable | 模型 model |
---|---|---|
幂函数 power function | D | W=aDb |
DH | W=a(DH)b | |
D2H | W=a(D2H)b | |
多项式函数 polynomial function | D | W=a+bD+cD2 |
D2 | W=a+bD2+cD4 | |
DH | W=a+bDH+c(DH)2 | |
D2H | W=a+bD2H+c(D2H)2 | |
指数函数 exponential function | D | W=aebD |
D2 | W=a | |
DH | W=aebDH | |
D2H | W=a |
Table 4
Comparisons of evaluation indexes of different independent variables"
自变量 independent variable | 单株single bamboo | 秆 stalk | 枝 branch | 叶 leaf |
---|---|---|---|---|
D | 0.845~0.950 | 0.834~0.945 | 0.750~0.832 | 0.567~0.724 |
D2 | 0.843~0.924 | 0.837~0.917 | 0.657~0.760 | 0.570~0.646 |
DH | 0.818~0.973 | 0.807~0.975 | 0.701~0.821 | 0.546~0.714 |
D2H | 0.660~0.971 | 0.647~0.971 | 0.574~0.831 | 0.480~0.722 |
Table 5
The biomass model of single bamboo and various organs aboveground"
竹龄/a bamboo age | 器官 organ | 函数类型 model type | 自变量 independent variable | 方程 fitting equation | R2 | RMSE/kg | MAE/kg |
---|---|---|---|---|---|---|---|
全竹龄 whole age classes | 单株 | 幂函数 | DH | W=0.11 4(DH)1.411 1 | 0.973** | 0.177 | 0.136 |
单株 | 幂函数 | D | W=0.107 0D2.334 1 | 0.950** | 0.240 | 0.185 | |
竹秆 | 幂函数 | DH | W=0.007 6(DH)1.489 1 | 0.975** | 0.180 | 0.141 | |
竹秆 | 幂函数 | D | W=0.081 1D2.450 0 | 0.946** | 0.264 | 0.202 | |
竹枝 | 幂函数 | D | W=0.015 2D-4.188 5 | 0.832** | 0.355 | 0.266 | |
竹叶 | 幂函数 | D | W=0.008 8D1.545 7 | 0.724** | 0.429 | 0.355 | |
1 | 单株 | 幂函数 | D2H | W=0.032 9D2H0.846 7 | 0.954** | 0.233 | 0.180 |
单株 | 幂函数 | D | W=0.114 8D2.308 3 | 0.945** | 0.253 | 0.199 | |
竹秆 | 幂函数 | D2H | W=0.021 1D2H0.909 1 | 0.965** | 0.217 | 0.174 | |
竹秆 | 幂函数 | D | W=0.081 8D2.467 1 | 0.948** | 0.264 | 0.203 | |
竹枝 | 幂函数 | D | W=0.014 5D1.827 2 | 0.833** | 0.368 | 0.273 | |
竹叶 | 幂函数 | D | W=0.007 9D1.654 3 | 0.769** | 0.408 | 0.331 | |
2 | 单株 | 多项式 | D2H | W=0.199 5+0.016 2D2H+ 0.000 004(D2H2) | 0.955** | 0.583 | 0.422 |
单株 | 幂函数 | D2 | W=2.334 4D0.120 0 | 0.944** | 0.181 | 0.138 | |
竹秆 | 多项式 | D2H | W=0.191 6+0.012 9D2H+ 0.000 000 2(D2H)2 | 0.964** | 0.456 | 0.322 | |
竹秆 | 幂函数 | D2 | W=2.415 1D0.088 9 | 0.938** | 0.198 | 0.153 | |
竹枝 | 幂函数 | D | W=0.016 7D2.183 | 0.774** | 0.365 | 0.290 | |
竹叶 | 多项式 | DH | W=0.115 8+0.003 9DH+ 0.000 08(DH)2 | 0.863** | 0.056 | 0.043 | |
竹叶 | 幂函数 | D | W=2.438D0.004 8 | 0.807** | 0.370 | 0.298 | |
3 | 单株 | 幂函数 | DH | W=0.027 6DH1.258 7 | 0.951** | 0.220 | 0.153 |
单株 | 幂函数 | D | W=0.124 3D2.366 3 | 0.932** | 0.259 | 0.169 | |
竹秆 | 幂函数 | DH | W=0.017 5DH1.320 2 | 0.964** | 0.196 | 0.137 | |
竹秆 | 幂函数 | D | W=0.056D2.475 5 | 0.940** | 0.253 | 0.178 | |
竹枝 | 幂函数 | D | W=0.349 6D1.786 2 | 0.814** | 0.353 | 0.294 | |
竹叶 | 指数函数 | D | W=0.009 1e0.629 1D | 0.791** | 0.472 | 0.387 |
Table 6
Comparison of the fitting effect between whole age classes and different age classes"
竹龄 bamboo age | 器官organ | R2 | RMSE/kg | MAE/kg |
---|---|---|---|---|
全竹龄 whole age | 单株single bamboo | 0.660~0.973 | 0.177~0.627 | 0.136~0.484 |
竹秆stalk | 0.647~0.975 | 0.180~0.673 | 0.141~0.518 | |
竹枝branch | 0.574~0.832 | 0.057~0.565 | 0.043~0.411 | |
竹叶leaf | 0.480~0.724 | 0.033~0.588 | 0.026~0.492 | |
不同竹龄 different age | 单株single bamboo | 0.654~0.955 | 0.233~0.769 | 0.180~0.589 |
竹秆stalk | 0.644~0.965 | 0.217~0.688 | 0.169~0.544 | |
竹枝branch | 0.633~0.833 | 0.173~0.601 | 0.133~0.440 | |
竹叶leaf | 0.535~0.863 | 0.027~0.444 | 0.021~0.295 |
Table 7
Biomass allocation situation of stand in different age classes"
竹龄/a bamboo age | 立竹密度/ (株·hm-2) quantity | 林分生物量/(t·hm-2) biomass | |||
---|---|---|---|---|---|
秆 stalk | 枝 branch | 叶 leaf | 总生物量 total biomass | ||
1 | 2 930±456 b | 11.408±2.274 b (91.61、33.68) | 0.740±0.167 b (5.94、19.71) | 0.305±0.064 c (2.45、22.10) | 12.560±2.505 b (100、31.92) |
2 | 3 641±580 a | 15.658±5.602 a (85.13、46.23) | 2.017±0.721 a (10.97、53.69) | 0.717±0.256 a (3.90、51.96) | 18.407±6.580 a (100、47.15) |
3 | 1 681±334 c | 6.805±1.552 c (83.36、20.09) | 1.000±0.354 c (12.25、26.62) | 0.358±0.154 b (4.39、25.94) | 8.163±2.450 c (100、20.93) |
总量 | 8 252±1 488 | 33.871±8.353 (86.83、100) | 3.757±0.909 (9.63、100) | 1.380±0.332 (3.54、100) | 39.008±9.593 (100、100) |
Table 8
Biomass allocation situation of stand in different diameter classes"
径级/cm diameter | 竹林密度/ (株·hm-2) density | 平均株高/m mean height | 林分生物量/(t·hm-2) stand biomass | |||
---|---|---|---|---|---|---|
秆 stalk | 枝 branch | 叶 leaf | 总生物量 total biomass | |||
1.6~2.5 | 111±47 e | 6.40 e | 0.047±0.024 d (80.73 b、0.14 d) | 0.008±0.003 c (14.04 a、0.21 c) | 0.003±0.001 c (2.74 b、0.22 c) | 0.058±0.028 c (100、0.15 c) |
2.6~3.5 | 314±88 d | 10.08±2.35 d | 0.442±0.118 c (85.75 a、1.30 c) | 0.054±0.017 c (10.29 b、1.44 c) | 0.021±0.005 c (3.95 a、1.52 c) | 0.517±0.140 c (100、1.33 c) |
3.6~4.5 | 1 948±616 b | 12.21±2.89 c | 5.402±1.830 b (86.19 a、15.95 b) | 0.635±0.212 b (10.15 b、16.90 b) | 0.228±0.074 b (3.65 a、16.52 b) | 6.265±2.116 b (100、16.06 b) |
4.6~5.5 | 4 589±1 742 a | 13.73±3.44 b | 21.265±6.237 a (87.10 a、62.78 a) | 2.311±0.708 a 9.45 b、61.51 a | 0.844±0.254 a (3.45 a、61.16 a) | 24.420±7.195 a (100、62.60 a) |
5.6~6.5 | 1 081±333 c | 15.07±4.10 a | 6.715±2.209 b (86.61 a、19.83 b) | 0.749±0.233 b (9.73 b、19.94 b) | 0.284±0.099 b (3.66 a、20.58 b) | 7.748±2.537 b (100、19.86 b) |
总量 | 8 043±1 827 | 12.02±3.12 | 33.871±8.353 (86.83、100) | 3.757±0.909 (9.63、100) | 1.380±0.332 (3.54、100) | 39.008±9.593 (100、100) |
[1] | 中国科学院中国植物志编辑委员会. 中国植物志:第九卷第一分册[M]. 北京: 科学出版社, 1996. |
[2] | 汪淑芳, 梁梓, 杨瑶君, 等. 不同秆龄慈竹竹秆含水率与生物量的分配特征及其生物量模型的构建[J]. 西部林业科学, 2013,42(1):42-45. |
WANG S F, LIANG Z, YANG Y J, et al. Biomass distribution characteristics and model construction of Neosinocalamus affinis at different ages[J]. J West China For Sci, 2013,42(1):42-45. DOI: 10.3969/j.issn.1672-8246.2013.01.008. | |
[3] | 唐建维, 萧自位, 王小李, 等. 西双版纳不同林龄人工龙竹林的生物量及其模型[J]. 中南林业科技大学学报, 2011,31(12):54-60. |
TANG J W, XIAO Z W, WANG X L, et al. Biomass and its regression models of artificial Dendrocalamus giganteus within different stand ages in Xishuangbanna[J]. J Central South Univ For Technol, 2011,31(12):54-60. DOI: 10.3969/j.issn.1673-923X.2011.12.011. | |
[4] | 徐道旺, 陈少红, 杨金满. 毛环竹笋用林生物量结构调查分析[J]. 福建林业科技, 2004,31(1):67-70. |
XU D W, CHEN S H, YANG J M. Investigation and analysis on the biomass structure of Phyllostachys meyeri woods for bamboo shoot[J]. J Fujian For Sci Technol, 2004,31(1):67-70. DOI: 10.3969/j.issn.1002-7351.2004.01.018. | |
[5] | 王海霞, 彭九生, 曾庆南. 淡竹立竹生物量计算模型构建[J]. 世界竹藤通讯, 2014,12(6):6-10. |
WANG H X, PENG J S, ZENG Q N. Construction of biomass model for stand of Phylostachys glauca[J]. World Bamboo Rattan, 2014,12(6):6-10. DOI: 10.3969/j.issn.1672-0431.2014.06.002. | |
[6] | 苏文会, 顾小平, 官凤英, 等. 大木竹种群生物量结构及其回归模型[J]. 南京林业大学学报(自然科学版), 2006,30(5):51-54. |
SU W H, GU X P, GUAN F Y, et al. Biomass structure and its regression models of Bambusa wenchouensis population[J]. J Nanjing For Univ (Nat Sci Ed), 2006,30(5):51-54. DOI: 10.3969/j.issn.1000-2006.2006.05.012. | |
[7] | 鹿士杨, 潘丽萍, 彭晚霞, 等. 不同竹类人工林生态系统生物量空间分配格局[J]. 生态科学, 2018,37(4):123-129. |
LU S Y, PAN L P, PENG W X, et al. Characteristics of biomass allocation in different bamboo plantations[J]. Ecol Sci, 2018,37(4):123-129. DOI: 10.14108/j.cnki.1008-8873.2018.04.015. | |
[8] | 王曙光, 普晓兰, 丁雨龙, 等. 云南箭竹地上部分生物量模型研究[J]. 南京林业大学学报(自然科学版), 2010,34(1):141-144. |
WANG S G, PU X L, DING Y L, et al. A study on the aboveground biomass models of Fargesia yunnanensis[J]. J Nanjing For Univ (Nat Sci Ed), 2010,34(1):141-144. DOI: 10.3969/j.issn.1000-2006.2010.01.032. | |
[9] | 杨清, 苏光荣, 段柱标, 等. 版纳甜龙竹种群生物量结构及其回归模型[J]. 西北农林科技大学学报(自然科学版), 2008,36(7):127-134. |
YANG Q, SU G R, DUAN Z B, et al. Biomass structure and its regression models of Dendrocalamus hamiltonii Nees et Arn.ex. Munro population[J]. J Northwest A&f Univ (Nat Sci Ed), 2008,36(7):127-134. | |
[10] | 夏载深, 李秋枫. 硬头黄竹生长规律观察研究[J]. 重庆林业科技, 1991(2):12-18. |
XIA Z S, LI Q F. Study on the growth rule of Bambusa rigida[J]. J Chongqing For Sci Technol, 1991(2):12-18. | |
[11] | 周益权, 顾小平, 吴晓丽, 等. 丛生竹秆基各笋目的出笋成竹生物学特性研究[J]. 林业科学研究, 2011,24(3):314-320. |
ZHOU Y Q, GU X P, WU X L, et al. Study on biological characteristics of bamboo shooting from base of sympodial bamboo buds[J]. For Res, 2011,24(3):314-320. | |
[12] | 郑郁善, 梁鸿燊. 绿竹生物量模型研究[J]. 竹子研究汇刊, 1997,16(4):43-46. |
ZHENG Y S, LIANG H S. Studies on the models for biomass of Sinocalamue oldhami[J]. J Bamboo Res, 1997,16(4):43-46. | |
[13] | 张宇, 岳祥华, 漆良华, 等. 利用异速生长关系和地统计方法估算武夷山南麓毛竹林生物量[J]. 生态学杂志, 2016,35(7):1957-1962. |
ZHANG Y, YUE X H, QI L H, et al. Estimation of Phyllostachys edulis forest biomass in southern Wuyishan Mountain using allometric equation and geostatistical technique[J]. Chin J Ecol, 2016,35(7):1957-1962. DOI: 10.13292/j.1000-4890.201607.036. | |
[14] | 黎曦. 赣南毛竹、硬头黄竹、坭竹等竹林生物量的研究[D]. 南京:南京林业大学, 2007. |
[15] | 张鹏, 黄玲玲, 张旭东, 等. 滩地硬头黄竹生物量结构及回归模型的研究[J]. 竹子研究汇刊, 2009,28(3):25-28. |
ZHANG P, HUANG L L, ZHANG X D, et al. Studies on biomass structure and its regression models of Bambusa rigida Keng[J]. J Bamboo Res, 2009,28(3):25-28. DOI: 10.3969/j.issn.1000-6567.2009.03.006. | |
[16] | 曹明勇. 长宁县竹林生物量及初级生产力特征研究[D]. 长沙:中南林业科技大学, 2016. |
CAO M Y. Study on bamboo biomass and primary productivity features with Changning County[D]. Changsha:Central South University of Forestry & Technology, 2016. | |
[17] | 陈霖. 宜宾市3个竹种地上部分生物量表的编制[D]. 雅安:四川农业大学, 2015. |
CHEN L. Establishment of aboveground biomass table of three bamboo species in Yibin[D]. Ya’an:Sichuan Agricultural University, 2015. | |
[18] | 李燕, 张建国, 段爱国, 等. 杉木人工林生物量估算模型的选择[J]. 应用生态学报, 2010,21(12):3036-3046. |
LI Y, ZHANG J G, DUAN A G, et al. Selection of biomass estimation models for Chinese fir plantation[J]. Chin J Appl Ecol, 2010,21(12):3036-3046. | |
[19] | 王太鑫, 丁雨龙, 李继清, 等. 巴山木竹种群生物量结构研究[J]. 竹子研究汇刊, 2005,24(1):19-24. |
WANG T X, DING Y L, LI J Q, et al. Studies on the structure of biomass of Arundianaria fargesii clone population[J]. J Bamboo Res, 2005,24(1):19-24. DOI: 10.3969/j.issn.1000-6567.2005.01.004. | |
[20] | 王维芳, 董薪明, 董小枫, 等. 森林生物量的空间自相关性研究[J]. 森林工程, 2018,34(2):35-39. |
WANG W F, DONG X M, DONG X F, et al. Study on spatial autocorrelation of forest biomass[J]. Forest Engineering, 2018,34(2):35-39. | |
[21] | 朱强根, 金爱武, 陈操, 等. 毛竹克隆构型及其生物量对不同营林模式的响应[J]. 南京林业大学学报(自然科学版), 2015,39(1):73-78. |
ZHU Q G, JIN A W, CHEN C, et al. The clonal architecture and biomass of Phyllostachys edulis as affected by different management modes[J]. J Nanjing For Univ (Nat Sci Ed), 2015,39(1):73-78. DOI: 10.3969/j.issn.1000-2006.2015.01.001. |
[1] | LI Minghui, LI Yuanfa. Evaluating influences of diameter classes on the species diversity of natural forests in the Nanpan River basin [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2023, 47(4): 166-174. |
[2] | XU Shanshan, LYU Jingyan, CHEN Fangyuan. Remote sensing vegetation detection method based on the deep convolutional neural network [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2022, 46(4): 185-193. |
[3] | SUN Caowen, ZHONG Wenwen, FU Xiangxiang, SHANG Xulan, FANG Shengzuo. A study on growth and aboveground biomass production of juvenile Cyclocarya paliurus plantations [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2022, 46(1): 138-144. |
[4] | YAO Wenjing, WANG Ru, LIN Shuyan, WANG Xing, YANG Meng, ZHENG Yi, DING Yulong. Growth mechanisms and model fitting of module biomass of Pleioblastus pygmaeus seedlings [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2020, 44(6): 103-110. |
[5] | HAN Yuna, ZHANG Yu, JIN Guangze. Effects of decay class and diameter class on moisture content and wood density in a typical mixed broadleaf-Korean pine forest [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2020, 44(2): 133-140. |
[6] | XU Baokun, XU Xiaogang,LI Yao, LI Xiaodong, CHEN Shuifei,DING Hui, JIANG Xiaohui, GOU Lingchen,FANG Yanming. Interspecific association analysis of Castanopsis eyrei community in evergreen broad-leaved forests in Huangshan, Anhui Province [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2019, 43(04): 77-84. |
[7] | ZHANG Xijin, LENG Hanbing, ZHAO Guangqi, JING Jun, TU Aicui, SONG Kun1,4*, DA Liangjun1,3,4. Allometric models for estimating aboveground biomass for four common greening tree species in Shanghai City, China [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2018, 42(02): 141-146. |
[8] | DENG Lin, ZHAN Hui, HE Wenzhi, ZHAO Jingwei, WANG Shuguang. Developmental changes of fiber wall of Fargesia fungosa [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2017, 41(04): 147-152. |
[9] | OU Xiaolan, LIU Yanhong. Effect of age, slope aspects and diameter classes on leaf functional traits of Pinus tabulaeformis in Songshan, Beijing [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2017, 41(04): 80-88. |
[10] | ZHU Yali, ZHANG Huifang,GAO Yaqi, DILIXIATI·Baoerhan, DING Chengfeng, YANG Yiyuan. Analysis on carbon content factors of Larix sibirica Ledeb. in Xinjiang [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2017, 41(03): 198-202. |
[11] | ZHANG Huifang, ZHU Yali, DILIXIATI·Baoerhan, GAO Yaqi, DING Chengfeng, WANG Lei. Biomass allocation patterns of Picea obovata and Larix sibirica in the Altai Mountains forest area [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2017, 41(01): 203-208. |
[12] | ZENG Wei, JIANG Bin, YU Lin, XIAO Fuming, XU Haining. Biomass allocation and its model construction of Cunninghamia lanceolata plantations in Jiangxi Province [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2016, 40(03): 177-182. |
[13] | HE Lieyan, KANG Xingang*, FAN Xiaoli, GAO Yan*. Estimation and analysis of understory shrub biomass in Changbai Mountains [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2011, 35(05): 45-50. |
[14] | LU Qiuling, ZHENG Abao, RUAN Honghuan. The carbon density and carbon storage of forests in Wawushan Forest Farm [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2010, 34(05): 115-119. |
[15] | Author Song DingquanJiang Zhilin (Nanjing Forestry UniversityNanjing210037) Zheng Zuomeng. A Preliminary Study on Population Natural Thinning Adjustment of Betula Luminifera [J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 1999, 23(03): 24-27. |
Viewed | ||||||
Full text |
|
|||||
Abstract |
|
|||||