Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances

doi:10.12302/j.issn.1000-2006.202104044

JOURNAL OF NANJING FORESTRY UNIVERSITY ›› 2022, Vol. 46 ›› Issue (3): 203-212.doi: 10.12302/j.issn.1000-2006.202104044

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Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances

HU Xingfeng¹(), WU Fan¹, SUN Xiaobo¹, CHEN Houping², YIN Anzheng², JI Kongshu¹^,^*()

1. Key Laboratory of Forest Genetics & Biotechnology of Ministry of Education, Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing 210037, China
2. State-owned Forest Tree Breeding Farm, Yu’an District, Lu’an City, Anhui Province, Lu’an 237010, China

Received:2021-04-26 Accepted:2021-11-08 Online:2022-05-30 Published:2022-06-10
Contact: JI Kongshu E-mail:xingfenghu@126.com;ksji@njfu.edu.cn

Abstract

Abstract:

【Objective】The growth provenance and wood property variation of mature masson pine (Pinus massoniana) are studied, revealing the variation rules for different traits at the level of geographical provenance, and screening suitable provenances for Lu’an City in Anhui Province and similar geographical areas.【Method】We selected the state-owned forest tree seed field in Yu’an District, Lu’an City, Anhui Province as the experimental site. 55 provenances were selected for 38-year-old (mature forest) P. massoniana as the test material. The tree height, diameter at breast height(DBH) volume, length and width of tracheid, basic density and microfibril angle are combined to analyze the growth and material properties.【Result】It was found that the diameter at breast height, tree height, volume, tracheid length, tracheid width, microfibril angle and basic density all showed highly significant differences at the provenance level (P < 0.01). The variation range of the phenotypic coefficients of variation and the genetic variation coefficients of each trait are 3.02%-58.61% and 5.56%-28.16%, respectively. The provenance heritability of the DBH, tree height, volume, tracheid length, tracheid width, basic density and microfibril angle are 0.665 7, 0.558 9, 0.616 9, 0.996 3, 0.965 6, 0.739 3 and 0.823 3, respectively. Growth traits are significantly negatively correlated with latitude, and are not significantly different from longitude. Wood properties at different latitude and longitude are both insignificantly different, but show local variation. In the correlation study, it was found that the correlation between growth traits is higher than that between growth and wood properties. The microfibril angle from the medulla core to the bark decreases with increase in age. The basic wood density, tracheid length and width from the pith to the bark increases with the gradual increase in age class, and the range of change decreases with the increase in age class. Through the comprehensive index, five provenances were selected with excellent comprehensive traits. The average genetic gains of DBH, tree height, volume, tracheid length and width, basic density and microfibril angle were 3.69%, 5.72%, 15.12%, 7.05%,-1.12%, 0.17% and -6.46%.【Conclusion】There is abundant genetic variation among the provenances of P. massoniana in mature forests, and growth characteristics show a variation law that gradually decreases from south to north, with wood properties not being related to latitude and longitude. The joint selection of pulp material growth and material properties provides a reliable basis for the selection of fine varieties of pulp material in P. massoniana.

Key words: masson pine (Pinus massoniana), provenance, basic density, tracheid, microfibril angle, wood property

CLC Number:

S722

HU Xingfeng, WU Fan, SUN Xiaobo, CHEN Houping, YIN Anzheng, JI Kongshu. Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances[J]. JOURNAL OF NANJING FORESTRY UNIVERSITY, 2022, 46(3): 203-212.

Figures/Tables 11

Table 1

Numbered list of sampling provenances of Pinus massoniana"

种源编号 provenance No.	种源 provenance	东经/(°) longitude (E)	北纬(°) latitude(N)	种源编号 provenance No.	种源 provenance	东经/(°) longitude (E)	北纬(°) latitude(N)
1	陕西城固	107.353 3	33.157 1	33	贵州黎平	109.140 1	26.237 4
2	河南新县	114.891 8	31.650 2	34	四川蒲江	103.509 1	30.203 0
3	安徽霍山	116.337 2	31.399 6	35	浙江鄞县	121.553 1	29.823 7
4	浙江遂昌	119.278 4	28.600 0	36	江西崇义	114.313 4	25.686 6
5	江西安远	115.394 2	25.142 7	39	江西崇仁	116.081 8	27.757 5
6	湖南慈利	111.148 9	29.434 5	40	浙江三门	121.401 4	29.112 9
7	湖北通山	114.487 7	29.613 6	41	安徽太平	118.143 3	30.299 6
8	福建漳平	117.427 1	25.296 3	42	浙江开化	118.420 9	29.143 2
9	广东高川	110.853 7	21.923 0	43	江西石城	116.346 3	26.320 0
11	贵州黄平	107.919 5	26.911 3	44	福建古田	118.746 8	26.581 7
12	四川南江	106.831 0	32.353 0	45	广东信宜	110.952 4	22.360 6
13	湖南安化	111.217 8	28.380 6	46	广西忻城	108.671 4	24.071 4
14	浙江庆远	119.066 0	27.626 0	47	浙江嵊县	120.837 4	29.567 3
15	江西安福	114.622 5	27.399 6	48	江西万载	114.453 7	28.114 7
17	湖南临湘	113.458 0	29.482 8	49	陕西南郑	106.940 1	33.006 1
18	福建德化	118.240 6	25.498 9	50	河南鹤峰	114.303 2	35.754 0
19	广东广宁	112.445 1	23.642 6	51	安徽屯溪	118.320 3	29.703 1
20	贵州都匀	107.531 1	26.267 7	52	浙江民生	120.752 9	30.104 0
21	浙江缙云	120.091 7	28.665 7	53	江西清江	114.798 0	29.193 5
22	江西吉安	114.997 6	27.120 2	54	湖南资兴	113.239 1	25.982 4
25	贵州凯里	107.986 3	26.574 1	55	福建邵武	117.501 1	27.348 5
26	河南固始	115.658 8	32.174 7	58	广西茶城	110.251 2	24.490 7
27	安徽马头	118.509 3	30.788 3	59	贵州德江	108.124 1	28.272 5
28	浙江永康	120.052 8	28.894 2	60	四川涪陵	107.394 7	29.707 8
29	江西余江	116.860 2	28.206 2	61	湖南永顺	109.862 4	28.987 8
30	湖南江永	111.345 8	25.282 3	63	江西资溪	109.862 4	28.987 8
31	福建三明	117.648 9	26.270 3	64	浙江西溪	120.080 0	30.271 9
32	广东罗定	111.573 0	22.776 4

Table 1

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性状 trait	均值 mean	标准差 SD	表型变异系数/% phenotypic coefficient of variation
DBH/cm	19.40	5.305 8	27.49
H/m	16.470 1	2.380 0	14.54
V/m³	0.251 1	0.144 0	58.61
TL/μm	3285.60	99.773 4	3.02
TW/μm	43.87	2.073 7	4.95
BD/(g·cm^-3)	0.568 9	0.033 9	5.93
MFA/(°)	29.23	1.654 7	5.70

性状 trait	变异来源 source of variation	自由度 df	均方 MS	F	P	显著性 signi- ficance
DBH	区组效应	5	418.930 6	30.366 4	1.82×10^-24	**
	种源效应	54	41.264 3	2.991 1	2.64×10⁹	**
H	区组效应	5	48.503 1	13.739 2	5.85×10^-12	**
	种源效应	54	8.004 2	2.267 3	9.92×10^-6	**
V	区组效应	5	0.243 5	19.966 7	6.18×10^-17	**
	种源效应	54	0.031 8	2.610 4	2.12×10^-7	**
TL	区组效应	5	48 078.271 4	24.986 3	1.10×10	**
	种源效应	54	525 894.681 6	273.307 2	9.81×10^-207	**
TW	区组效应	5	1.474 1	1.118 9	3.51×10^-1
	种源效应	54	38.298 2	29.070 7	3.60×10^-85	**
BD	区组效应	5	0.002 6	1.949 1	5.14×10^-2
	种源效应	54	0.005 2	3.836 4	2.64×10^-15	**
MFA	区组效应	5	18.105 9	5.407 1	5.78×10³	**
	种源效应	54	18.952 9	5.660 0	1.13×10^-14	**

性状 trait	方差variance						变异系数/% CV			种源遗传力 provenance heritability
性状 trait	表型 phenotypic	环境 environmental		遗传 genetic		表型 phenotypic		遗传 genetic		种源遗传力 provenance heritability
DBH	28.151 5		9.7090		1.381 0		27.49		7.13	0.665 7
H	5.664 4		0.8680		6.282 0		14.54		12.92	0.558 9
V	0.020 7		0.005 0		0.005 0		58.61		28.16	0.616 9
TL	9 954.731 3		874.155 0		87 648.621 0		3.02		9.01	0.996 3
TW	4.725 0		0.025 0		6.392 0		4.95		5.76	0.965 6
BD	0.001 1		0.000 048		0.001 0		5.93		5.56	0.739 3
MFA	2.738 0		0.329 0		6.336 0		5.70		8.60	0.823 3

项目item	H	DBH	V	TL	TW	BD	MFA
经度longitude(E)	-0.014(0.920)	-0.070(0.612)	-0.036(0.794)	-0.006(0.967)	-0.040(0.770)	0.094(0.494)	-0.072(0.599)
纬度latitude(N)	-0.427^**(0.001)	-0.561^**(<0.01)	-0.539^**(<0.01)	-0.071(0.606)	-0.140(0.308)	0.208(0.128)	0.011(0.939)

性状 trait	自由度 df	均方 MS	F	P	显著性 sig
TL	7	4 322.129 0	313.344 0	1.08×10^-274	***
TW	7	3.746 0	825.129 0	P<0.01	***
BD	7	1 012 684 377.190 0	6 467.518 0	P<0.01	***
MFA	7	16 358.522 0	391.205 0	P<0.01	***

Joint analysis of growth and wood property of 38-year-old Pinus massoniana from 55 provenances

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性状trait	DBH	H	V	TL	TW	BD	MFA
DBH	—	0.635^**	0.963^**	-0.016	0.018	-0.231^**	0.010
		(<0.01)	(<0.01)	(0.624)	(0.575)	(<0.01)	(0.899)
H	0.632^**	—	0.719^**	-0.008	0.023	-0.055	0.024
	(<0.01)		(<0.01)	(0.794)	(0.477)	(0.225)	(0.758)
V	0.735^**	0.967^**	—	0	0.025	-0.197^**	-0.008
	(<0.01)	(<0.01)		(0.994)	(0.438)	(<0.01)	(0.915)
TL	0.047	0.091	0.124	—	0.375^**	0.054	-0.098
	(0.733)	(0.507)	(0.369)		(<0.01)	(0.229)	(0.211)
TW	0.203	0.147	0.204	0.524^**	—	-0.029	0.027
	(0.137)	(0.284)	(0.135)	(<0.01)		(0.520)	(0.734)
BD	-0.417^**	-0.581^**	-0.516^**	0.120	-0.087	—	0.036
	(0.002)	(<0.01)	(<0.01)	(0.385)	(0.526)		(0.645)
MFA	-0.333^*	-0.121	-0.155	-0.098	-0.028	0.033	—
	(0.013)	(0.377)	(0.259)	(0.478)	(0.839)	(0.811)

种源号 provenance No.	H/m	DBH/cm	V/m³	TL/μm	TW/μm	BD/(g·cm^-3)	MFA/(°)	I
5	18.93	22.01	0.355 1	3 644.5	46.9	0.601 6	24.8	9.342
8	17.28	22.11	0.344 2	3 399.7	42.9	0.591 6	29.4	8.042
13	17.96	19.93	0.286 9	3 470.8	42.3	0.558 4	24.8	8.584
40	16.18	18.78	0.233 4	3 403.9	41.8	0.569 8	24.7	8.057
58	16.55	24.10	0.343 6	3 672.5	42.9	0.529 6	31.1	7.753
总体均值 population mean	16.47	19.40	0.251 1	3 285.6	43.9	0.568 9	29.3	—
入选群体均值 selected group mean	17.38	21.39	0.312 7	3 518.3	43.4	0.570 2	27.0	—
遗传增益/% genetic gain	3.69	5.72	15.12	7.05	-1.12	0.17	-6.46	—

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